Rewilded horses in European nature conservation – a genetics, ethics, and welfare perspective.

1. Ajmone‐Marsan P, Garcia JF, Lenstra JA. On the origin of cattle: how aurochs became cattle and colonized the world. Evolutionary Anthropology 19, 148–157.
2. Al Jassim RAM, Andrews FM. The bacterial community of the horse gastrointestinal tract and its relation to fermentative acidosis, laminitis, colic, and stomach ulcers. The Veterinary Clinics of North America. Equine Practice 25, 199–215.
3. Alessandri G, Milani C, Mancabelli L, Mangifesta M, Lugli GA, Viappiani A, Duranti S, Turroni F, Ossiprandi MC, van Sinderen D, Ventura M. The impact of human‐facilitated selection on the gut microbiota of domesticated mammals. FEMS Microbiology Ecology 95, 121.
4. Allendorf FW, Chris Funk W, Aitken SN, Byrne M, Luikart G. Conservation and the Genomics of Populations. Oxford University Press Oxford, 2012.
5. Andrews FM, Buchanan BR, Smith SH, Elliott SB, Saxton AM. In vitro effects of hydrochloric acid and various concentrations of acetic, propionic, butyric, or valeric acids on bioelectric properties of equine gastric squamous mucosa. American Journal of Veterinary Research 67, 1873–1882.
6. Andrieu J, Henry S, Hausberger M, Thierry B. Informed horses are influential in group movements, but they may avoid leading. Animal Cognition 19, 451–458.
7. Ang L, Vinderola G, Endo A, Kantanen J, Jingfeng C, Binetti A, Burns P, Qingmiao S, Suying D, Zujiang Y, Rios‐Covian D, Mantziari A, Beasley S, Gomez‐Gallego C, Gueimonde M. Gut microbiome characteristics in feral and domesticated horses from different geographic locations. Communications Biology 5, 172.
8. Anthony DW. The horse, the wheel and language. In How Bronze‐Age Riders from the Steppes Shaped the Modern World. Princeton University Press Princeton, NJ, 2007.
9. Anthony DW. The Sintashta genesis: the roles of climate change, warfare, and long‐distance trade. In Social Complexity in Prehistoric Eurasia: Monuments, Metals and Mobility. Cambridge University Press Cambridge, 2009.
10. Asplin KE, Sillence MN, Pollitt CC, McGowan CM. Induction of laminitis by prolonged hyperinsulinaemia in clinically normal ponies. Veterinary Journal 174, 530–535.
11. Bakker ES, Svenning J‐C. Trophic rewilding: impact on ecosystems under global change. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 373, 20170432.
12. Bangs E, Fontaine J, Jimenez M, Meier T, Bradley E, Niemeyer C, Smith DW, Mack CM, Asher VJ, Oakleaf JK. People and wildlife: managing wolf–human conflict in the northwestern United States. In People and Wildlife, Conflict or Co‐Existence?. Cambridge University Press Cambridge, 2005.
13. Banse K. Do we live in a largely top‐down regulated world?. Journal of Biosciences 32, 791–796.
14. Barmentlo NWG, Meirmans PG, Stiver WH, Yarkovich JG, McCann BE, Piaggio AJ, Wright D, Smyser TJ, Bosse M. Natural selection on feralization genes contributed to the invasive spread of wild pigs throughout the United States. Molecular Ecology 33, e17383.
15. Barth JMI, Handley SA, Kintzl D, Leonard G, Malinsky M, Matschiner M, Meyer BS, Salzburger W, Stefka J, Trucchi E. The history and organization of the workshop on population and speciation genomics. Evolution 16, 2.
16. Bechert US, Turner JW Jr, Baker DL, Eckery DC, Bruemmer JE, Lyman CC, Prado TM, King SRB, Fraker MA. Fertility control options for management of free‐roaming horse populations. Human‐Wildlife Interactions 16, 5.
17. Beeton NJ, Johnson CN. Modelling horse management in the Australian Alps. Ecological Management & Restoration 20, 57–62.
18. Beever EA, Herrick JE. Effects of feral horses in Great Basin landscapes on soils and ants: direct and indirect mechanisms. Journal of Arid Environments 66, 96–112.
19. Beever EA, Huntsinger L, Petersen SL. Conservation challenges emerging from free‐roaming horse management: a vexing social‐ecological mismatch. Biological Conservation 226, 321–328.
20. Behr DM, Ozgul A, Cozzi G. Combining human acceptance and habitat suitability in a unified socio‐ecological suitability model: a case study of the wolf in Switzerland. The Journal of Applied Ecology 54, 1919–1929.
21. Bekoff M. The importance of ethics in conservation biology: Let's be ethicists not ostriches. Endangered Species Update 19, 23–26.
22. Belaunzaran X, Bessa RJB, Lavín P, Mantecón AR, Kramer JKG, Aldai N. Horse‐meat for human consumption—current research and future opportunities. Meat Science 108, 74–81.
23. Bell RJW, Kingston JK, Mogg TD, Perkins NR. The prevalence of gastric ulceration in racehorses in New Zealand. New Zealand Veterinary Journal 55, 13–18.
24. Bellone RR, Holl H, Setaluri V, Devi S, Maddodi N, Archer S, Sandmeyer L, Ludwig A, Foerster D, Pruvost M, Reissmann M, Bortfeldt R, Adelson DL, Lim SL, Nelson J. Evidence for a retroviral insertion in TRPM1 as the cause of congenital stationary night blindness and leopard complex spotting in the horse. PLoS One 8, e78280.
25. Berger J. Reproductive fates of dispersers in a harem‐dwelling ungulate: the wild horse. In Mammalian Dispersal Patterns: The Effects of Social Structure on Population Genetics. University of Chicago Press Chicago, IL, 1987.
26. Bernáldez‐Sánchez E, García‐Viñas E, Sanguino F, Villalón D, Leonard JA. Equids (Equus sp.) in Southern Spain from the Palaeolithic to the Bronze Age. Journal of Quaternary Science 39, 261–276.
27. Bernatchez L, Wellenreuther M, Araneda C, Ashton DT, Barth JMI, Beacham TD, Maes GE, Martinsonh JT, Miller KM, Naish KA, Ovenden JR, Primmer CR, Young Suk H, Therkildsen NO, Withler RE. Harnessing the power of genomics to secure the future of seafood. Trends in Ecology & Evolution 32, 665–680.
28. Bhattacharyya J, Slocombe DS, Murphy SD. The ‘wild’ or ‘feral’ distraction: effects of cultural understandings on management controversy over free‐ranging horses (Equus ferus caballus). Human Ecology 39, 613–625.
29. Bickford D, Posa MRC, Qie L, Campos‐Arceiz A, Kudavidanage EP. Science communication for biodiversity conservation. Biological Conservation 151, 74–76.
30. Bonavent C, Olsen K, Ejrnæs R, Fløjgaard C, Hansen MD, Normand S, Svenning J‐C, Bruun HH. Grazing by semi‐feral cattle and horses supports plant species richness and uniqueness in grasslands. Applied Vegetation Science 26, 12718.
31. Borowska A, Wolska D, Niedzwiedz A, Borowicz H, Jaworski Z, Siemieniuch M, Szwaczkowski T. Some genetic and environmental effects on equine asthma in Polish Konik horses. Animals 11, 2285.
32. Boulbes N, van Asperen EN. Biostratigraphy and palaeoecology of European Equus. Frontiers in Ecology and Evolution 7, 301.
33. Bourgeon L, Burke A. Horse exploitation by Beringian hunters during the Last Glacial Maximum. Quaternary Science Reviews 269, 107140.
34. Bouskila A, Lourie E, Sommer S, de Vries H, Hermans ZM, van Dierendonck M. Similarity in sex and reproductive state, but not relatedness, influence the strength of association in the social network of feral horses in the Blauwe Kamer Nature Reserve. Israel Journal of Ecology & Evolution 61, 106–113.
35. Bowker RM. Contrasting structural morphologies of “good” and “bad” footed horses. American Association of Equine Practitioners Proceedings 49, 186–209.
36. Boyd L, Scorolli A, Nowzari H, Bouskila A. Wild equids: ecology, management, and conservation. In Social Organization of Wild Equids. Johns Hopkins University Press Baltimore, 2016.
37. Bracke MBM, Hopster H. Assessing the importance of natural behavior for animal welfare. Journal of Agricultural & Environmental Ethics 19, 77–89.
38. Bradshaw JWS, Casey RA. Anthropomorphism and anthropocentrism as influences in the quality of life of companion animals. In Animal Welfare. Cambridge University Press Cambridge, 2007.
39. Braunisch V, Home R, Pellet J, Arlettaz R. Conservation science relevant to action: a research agenda identified and prioritized by practitioners. Biological Conservation 153, 201–210.
40. Brosnahan MM, Brooks SA, Antczak DF. Equine clinical genomics: A clinician's primer. Equine Veterinary Journal 42, 658–670.
41. Brüniche‐Olsen A, Kellner KF, Anderson CJ, DeWoody JA. Runs of homozygosity have utility in mammalian conservation and evolutionary studies. Conservation Genetics 19, 1295–1307.
42. Callan R, Nibbelink NP, Rooney TP, Wiedenhoeft JE, Wydeven AP. Recolonizing wolves trigger a trophic cascade in Wisconsin (USA). The Journal of Ecology 101, 837–845.
43. Carroll SL, Sykes BW, Mills PC. Moving toward fear‐free husbandry and veterinary care for horses. Animals 12, 2907.
44. Carter GK, Maki JJ. Hoof cracks and wall defects. In Proceedings of the Annual Convention of the American Association of Equine Practitioners. Baltimore, 2010 .
45. Carver S, Convery I, Hawkins S, Beyers R, Eagle A, Kun Z, Van Maanen E, Cao Y, Fisher M, Edwards SR, Nelson C, Gann GD, Shurter S, Aguilar K, Andrade A. Guiding principles for rewilding. Conservation Biology 35, 1882–1893.
46. Chodkiewicz A. Advantages and disadvantages of polish primitive horse grazing on valuable nature areas – a review. Global Ecology and Conservation 21, e00879.
47. Christensen JW. Fear in Horses: Responses to Novelty and Habituation. Swedish University of Agricultural Sciences Uppsala, 2006.
48. Cieslak M, Pruvost M, Benecke N, Hofreiter M, Morales A, Reissmann M, Ludwig A. Origin and history of mitochondrial DNA lineages in domestic horses. PLoS One 5, e15311.
49. Cointreau M. Equine Herbs and Healing ‐ An Earth Lodge Pocket Guide to Holistic Horse Wellness. Earth Lodge Gaylordswill, CT, 2016.
50. Colpitts J, McLoughlin PD, Poissant J. Runs of homozygosity in Sable Island feral horses reveal the genomic consequences of inbreeding and divergence from domestic breeds. BMC Genomics 23, 501.
51. Cookson LJ. A definition for wildness. Ecopsychology 3, 187–193.
52. Corlett RT. The role of rewilding in landscape design for conservation. Current Landscape Ecology Reports 1, 127–133.
53. Cornelissen P. Large Herbivores as a Driving Force of Woodland‐Grassland Cycles: The Mutual Interactions between the Population Dynamics of Large Herbivores and Vegetation Development in a Eutrophic Wetland. Wageningen University and Research Wageningen, 2017.
54. Cornelissen P, Bokdam J, Sykora K, Berendse F. Effects of large herbivores on wood pasture dynamics in a European wetland system. Basic and Applied Ecology 15, 396–406.
55. Correa‐Fiz F, Blanco‐Fuertes M, Navas MJ, Lacasta A, Bishop RP, Githaka N, Onzere C, Le Potier MF, Almagro‐Delgado V, Martinez J, Aragon V. Comparative analysis of the fecal microbiota from different species of domesticated and wild suids. Scientific Reports 9, 13616.
56. Cosgrove EJ, Sadeghi R, Schlamp F, Holl HM, Moradi‐Shahrbabak M, Miraei‐Ashtiani SR, Abdalla S, Shykind B, Troedsson M, Stefaniuk‐Szmukier M, Prabhu A, Bucca S, Bugno‐Poniewierska M, Wallner B, Malek J. Genome diversity and the origin of the Arabian horse. Scientific Reports 10, 9702.
57. Cromsigt JPGM, Kuijper DPJ, Adam M, Beschta RL, Churski M, Eycott A, Kerley GIH, Mysterud A, Schmidt K, West K. Hunting for fear: innovating management of human–wildlife conflicts. Journal of Applied Ecology 50, 544–549.
58. Dalla Costa E, Minero M, Lebelt D, Stucke D, Canali E, Leach MC. Development of the Horse Grimace Scale (HGS) as a pain assessment tool in horses undergoing routine castration. PLoS One 9, e92281.
59. De Roos AM, Galic N, Heesterbeek H. How resource competition shapes individual life history for nonplastic growth: ungulates in seasonal food environments. Ecology 90, 945–960.
60. Dobbie WR, Dmck B, Braysher ML. Managing Vertebrate Pests: Feral Horses. Australian Government Publishing Service Canberra, 1993.
61. Driscoll DA, Worboys GL, Allan H, Banks SC, Beeton NJ, Cherubin RC, Doherty TS, Finlayson CM, Green K, Hartley R, Hope G, Johnson CN, Lintermans M, Mackey B, Paull DJ. Impacts of feral horses in the Australian Alps and evidence‐based solutions. Ecological Management & Restoration 20, 63–72.
62. Durant SM, Groom R, Kuloba B, Samna A, Muzuma U, Gadimang P, Mandisodza‐Chikerema R, Ipavec A, Mitchell N, Ikanda D, Msuha M. Bridging the divide between scientists and decision‐makers: how behavioural ecologists can increase the conservation impact of their research?. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 374, 20190011.
63. Durward‐Akhurst SA, Schaefer RJ, Grantham B, Carey WK, Mickelson JR, McCue ME. Genetic variation and the distribution of variant types in the horse. Frontiers in Genetics 12, 758366.
64. Engster D. Care ethics and animal welfare. Journal of Social Philosophy 37, 521–536.
65. Evans AR, Janis CM. The evolution of high dental complexity in the horse lineage. Annales Zoologici Fennici 51, 73–79.
66. Fages A, Hanghøj K, Khan N, Gaunitz C, Seguin‐Orlando A, Leonardi M, McCrory Constantz C, Gamba C, Al‐Rasheid KAS, Albizuri S, Alfarhan AH, Allentoft M, Alquraishi S, Anthony D, Baimukhanov N. Tracking five millennia of horse management with extensive ancient genome time series. Cell 177, 1419–1435.
67. Fages A, Seguin‐Orlando A, Germonpré M, Orlando L. Horse males became over‐represented in archaeological assemblages during the Bronze Age. Journal of Archaeological Science: Reports 31, 102364.
68. Feh C. Relationships and communication in socially natural horse herds. In The Domestic Horse: The Origins, Development and Management of its Behaviour. Cambridge University Press Cambridge, 2005.
69. Finno CJ, Spier SJ, Valberg SJ. Equine diseases caused by known genetic mutations. Veterinary Journal 179, 336–347.
70. Finno CJ, Stevens C, Young A, Affolter V, Joshi NA, Ramsay S, Bannasch DL. SERPINB11 frameshift variant associated with novel hoof specific phenotype in Connemara ponies. PLoS Genetics 11, e1005122.
71. Flauger B, Krueger K. Aggression level and enclosure size in horses (Equus caballus). Pferdeheilkunde 29, 495–504.
72. Fleurance G, Edouard N, Collas C, Duncan P, Farruggia A, Baumont R, Lecomte T, Dumont B. How do horses graze pastures and affect the diversity of grassland ecosystems? In Forages and Grazing in Horse Nutrition. Wageningen Academic Publishers Wageningen, 2012.
73. Fløjgaard C, De Barba M, Taberlet P, Ejrnæs R. Body condition, diet and ecosystem function of red deer (Cervus elaphus) in a fenced nature reserve. Global Ecology and Conservation 11, 312–323.
74. Fløjgaard C, Pedersen PBM, Sandom CJ, Svenning J‐C, Ejrnæs R. Exploring a natural baseline for large‐herbivore biomass in ecological restoration. Journal of Applied Ecology 59, 18–24.
75. Flood J, Stewart AJ. Non‐steroidal anti‐inflammatory drugs and associated toxicities in horses. Animals 12, 2939.
76. Florence L, McDonnell SM. Hoof growth and wear of semi‐feral ponies during an annual summer ‘self‐trimming’ period. Equine Veterinary Journal 38, 642–645.
77. Fornal A, Kowalska K, Zabek T, Piestrzynska‐Kajtoch A, Musiał AD, Ropka‐Molik K. Genetic diversity and population structure of Polish Konik horse based on individuals from all the male founder lines and microsatellite markers. Animals 10, 1569.
78. Forrest S. Inventing the wild horse: the manmade history of the Takhi and Tarpan from 1828 to 2018. In Horse Breeds and Human Society. Routledge Oxfordshire, 2019.
79. Frank N. Equine metabolic syndrome. The Veterinary Clinics of North America: Equine Practice 27, 73–92.
80. Frank N, Acvim D. Endocrinopathic laminitis, obesity‐associated laminitis, and pasture‐associated laminitis. In Proceedings of the Annual Convention of the American Association of Equine Practitioners. Baltimore, USA, 2008 .
81. Fraser MD, Stanley CR, Hegarty MJ. Recognising the potential role of native ponies in conservation management. Biological Conservation 235, 112–118.
82. Frey N, Beck JL, Singletary L, Snell L, Scasta D, Hadfield J. Western US residents' knowledge of wild free‐roaming horses and their management on federal public lands. Rangeland Ecology & Management 92, 12–23.
83. Frey SN, Scasta JD, Beck JL, Singletary L, Snell LK. Public knowledge of free‐roaming horses in the United States. Proceedings of the 30th Vertebrate Pest Conference, Paper 17 2022.
84. Frie AC. Horses and the embodiment of elite masculinity in the Dolenjska Hallstatt culture. Oxford Journal of Archaeology 37, 25–44.
85. Friedrich LA, Jefferson R, Glegg G. Public perceptions of sharks: gathering support for shark conservation. Marine Policy 47, 1–7.
86. Fuentes‐Pardo AP, Ruzzante DE. Whole‐genome sequencing approaches for conservation biology: advantages, limitations and practical recommendations. Molecular Ecology 26, 5369–5406.
87. Fulton GR, Ford HA. The conflict between animal welfare and conservation. Pacific Conservation Biology 7, 152–153.
88. Fureix C, Bourjade M, Henry S, Sankey C, Hausberger M. Exploring aggression regulation in managed groups of horses Equus caballus. Applied Animal Behaviour Science 138, 216–228.
89. Galinelli N, Wambacq W, Broeckx BJG, Hesta M. High intake of sugars and starch, low number of meals and low roughage intake are associated with Equine Gastric Ulcer Syndrome in a Belgian cohort. Journal of Animal Physiology and Animal Nutrition 105, 18–23.
90. Gamborg C, Gremmen B, Christiansen SB, Sandøe P. De‐domestication: ethics at the intersection of landscape restoration and animal welfare. Environmental Values 19, 57–78.
91. Gandiwa E. Top‐down and bottom‐up control of large herbivore populations: a review of natural and human‐induced influences. Tropical Conservation Science 6, 493–505.
92. Garber A, Hastie P, Murray J‐A. Factors influencing equine gut microbiota: current knowledge. Journal of Equine Veterinary Science 88, 102943.
93. Garrido P, Edenius L, Mikusiński G, Skarin A, Jansson A, Thulin C‐G. Experimental rewilding may restore abandoned wood‐pastures if policy allows. Ambio 50, 101–112.
94. Garrido P, Mårell A, Öckinger E, Skarin A, Jansson A, Thulin C‐G. Experimental rewilding enhances grassland functional composition and pollinator habitat use. The Journal of Applied Ecology 56, 946–955.
95. Garrido P, Naumov V, Söderquist L, Jansson A, Thulin C‐G. Effects of experimental rewilding on butterflies, bumblebees and grasshoppers. Journal of Insect Conservation 26, 763–771.
96. Gaunitz C, Fages A, Hanghøj K, Albrechtsen A, Khan N, Schubert M, Seguin‐Orlando A, Owens IJ, Felkel S, Bignon‐Lau O, de Barros Damgaard P, Mittnik A, Mohaseb AF, Davoudi H, Alquraishi S. Ancient genomes revisit the ancestry of domestic and Przewalski's horses. Science 360, 111–114.
97. Gavriliuc S, Reza S, Jeong C, Getachew F, McLoughlin PD, Poissant J. Targeted genome‐wide SNP genotyping in feral horses using non‐invasive fecal swabs. Conservation Genetics Resources 14, 203–213.
98. Gering E, Incorvaia D, Henriksen R, Conner J, Getty T, Wright D. Getting back to nature: feralization in animals and plants. Trends in Ecology & Evolution 34, 1137–1151.
99. Gerritsen V. Animal welfare in Switzerland – constitutional aim, social commitment, and a major challenge. Global Journal Animal Law 1, 2341–8168.
100. Giles SL, Nicol CJ, Rands SA, Harris PA. Assessing the seasonal prevalence and risk factors for nuchal crest adiposity in domestic horses and ponies using the Cresty Neck Score. BMC Veterinary Research 11, 13.
101. Giovos I, Barash A, Barone M, Barría C, Borme D, Brigaudeau C, Charitou A, Brito C, Currie J, Dornhege M, Endrizzi L, Forsberg K, Jung A, Kleitou P, MacDiarmid A. Understanding the public attitude towards sharks for improving their conservation. Marine Policy 134, 104811.
102. Goodwin D. Horse behaviour: evolution, domestication and feralisation. In The Welfare of Horses. Springer Netherlands Dordrecht, 2007.
103. Górecka‐Bruzda A, Jaworska J. The social and reproductive challenges faced by free‐roaming horse (Equus caballus) stallions. Animals 13, 1151.
104. Górecka‐Bruzda A, Jaworski Z, Jaworska J, Siemieniuch M. Welfare of free‐roaming horses: 70 years of experience with Konik Polski breeding in Poland. Animals 10, 1094.
105. Grange S, Duncan P, Gaillard J‐M. Poor horse traders: large mammals trade survival for reproduction during the process of feralization. Proceedings of the Royal Society. Biological sciences 276, 1911–1919.
106. Gruner DS. Attenuation of top‐down and bottom‐up forces in a complex terrestrial community. Ecology 85, 3010–3022.
107. Guest K, Mattfeld M. Purity, identity and the making of the modern horse. In Horse Breeds and Human Society. Routledge Oxfordshire, 2019.
108. Guthrie RD. Rapid body size decline in Alaskan Pleistocene horses before extinction. Nature 426, 169–171.
109. Haile J, Froese DG, Macphee RDE, Roberts RG, Arnold LJ, Reyes AV, Rasmussen M, Nielsen R, Brook BW, Robinson S, Demuro M, Gilbert MTP, Munch K, Austin JJ, Cooper A. Ancient DNA reveals late survival of mammoth and horse in interior Alaska. Proceedings of the National Academy of Sciences of the United States of America 106, 22352–22357.
110. Hampson BA, de Laat MA, Beausac C, Rovel T, Pollitt CC. Histopathological examination of chronic laminitis in Kaimanawa feral horses of New Zealand. New Zealand Veterinary Journal 60, 285–289.
111. Hampson BA, de Laat MA, Mills PC, Pollitt CC. Distances travelled by feral horses in ‘outback’ Australia. Equine Veterinary Journal 42, 582–586.
112. Hampson BA, de Laat MA, Mills PC, Walsh DM, Pollitt CC. The feral horse foot. Part B: radiographic, gross visual and histopathological parameters of foot health in 100 Australian feral horses. Australian Veterinary Journal 91, 23–30.
113. Hampson BA, Trezise A, Pollitt CC, Seddon J. The genetic analysis of Australian feral horses: the effect of environment on the feral horse foot. PhD thesis: School of Veterinary Science, The University of Queensland Australia, 2011.
114. Hampton JO, Edwards GP, Cowled BD, Forsyth DM, Hyndman TH, Perry AL, Miller CJ, Adams PJ, Collins T. Assessment of animal welfare for helicopter shooting of feral horses. Wildlife Research 44, 97–105.
115. Hampton JO, Hyndman TH, Barnes A, Collins T. Is wildlife fertility control always humane?. Animals 5, 1047–1071.
116. Hare D, Daniels M, Blossey B. Public perceptions of deer management in Scotland. Frontiers in Conservation Science 2, 781546.
117. Harrington LA, Moehrenschlager A, Gelling M, Atkinson RPD, Hughes J, Macdonald DW. Conflicting and complementary ethics of animal welfare considerations in reintroductions. Conservation Biology 27, 486–500.
118. Hartmann E, Søndergaard E, Keeling LJ. Keeping horses in groups: a review. Applied Animal Behaviour Science 136, 77–87.
119. Harvey AM, Ramp D, Mellor DJ. Review of the foundational knowledge required for assessing horse welfare. Animals 12, 3385.
120. Hausberger M, Roche H, Henry S, Visser EK. A review of the human–horse relationship. Applied Animal Behaviour Science 109, 1–24.
121. Henderson AJZ. Don't fence me in: managing psychological well being for elite performance horses. Journal of Applied Animal Welfare Science 10, 309–329.
122. Hennig JD, Duchardt CJ, Esmaeili S, Fuhlendorf SD, Beck JL, Francisco TI, Scasta JD. A crossroads in the rearview mirror: the state of United States feral equid management in 2023. Bioscience 73, 404–407.
123. Heuertz M, Carvalho SB, Galindo J, Rinkevich B, Robakowski P, Aavik T, Altinok I, Barth JMI, Cotrim H, Goessen R, González‐Martínez SC, Grebenc T, Hoban S, Kopatz A, McMahon BJ. The application gap: genomics for biodiversity and ecosystem service management. Biological Conservation 278, 109883.
124. Hewetson M, Tallon R. Equine squamous gastric disease: prevalence, impact and management. Veterinary Medicine 12, 381–399.
125. Hinchcliffe DL, Lea JMD, Palme R, Shultz S. Fecal glucocorticoid metabolites as biomarkers in equids: assay choice matters. The Journal of Wildlife Management 85, 1175–1186.
126. Hoban S, Bruford MW, Funk WC, Galbusera P, Griffith MP, Grueber CE, Heuertz M, Hunter ME, Hvilsom C, Stroil BK, Kershaw F, Khoury CK, Laikre L, Lopes‐Fernandes M, MacDonald AJ. Global commitments to conserving and monitoring genetic diversity are now necessary and feasible. Bioscience 71, 964–976.
127. Hoban SM, Hauffe HC, Pérez‐Espona S, Arntzen JW, Bertorelle G, Bryja J, Frith K, Gaggiotti OE, Galbusera P, Godoy JA, Hoelzel AR, Nichols RA, Primmer CR, Russo I‐R, Segelbacher G. Bringing genetic diversity to the forefront of conservation policy and management. Conservation Genetics Resources 5, 593–598.
128. Hongo A, Akimoto M. The role of incisors in selective grazing by cattle and horses. The Journal of Agricultural Science 140, 469–477.
129. Hopcraft JGC, Olff H, Sinclair ARE. Herbivores, resources and risks: alternating regulation along primary environmental gradients in savannas. Trends in Ecology & Evolution 25, 119–128.
130. Hulbert RC. Taxonomic evolution in North American Neogene horses (subfamily Equinae): the rise and fall of an adaptive radiation. Paleobiology 19, 216–234.
131. Hunter P. Genetics against extinction: new conservation strategies consider genetic diversity and habitat loss. EMBO Reports 24, e57521.
132. IUCN/SSC. Guidelines on the Use of Ex Situ Management for Species Conservation. Version 2.0. IUCN Species Survival Commission Gland, 2014.
133. Jagannathan V, Gerber V, Rieder S, Tetens J, Thaller G, Drögemüller C, Leeb T. Comprehensive characterization of horse genome variation by whole‐genome sequencing of 88 horses. Animal Genetics 50, 74–77.
134. Janis C. The evolutionary strategy of the Equidae and the origins of rumen and cecal digestion. Evolution 30, 757–774.
135. Jepson P, Schepers F. Making Space for Rewilding: Creating an Enabling Policy Environment. Rewilding Europe Oxford and Nijmegen, 2016.
136. Jepson P, Schepers F, Helmer W. Governing with nature: a European perspective on putting rewilding principles into practice. Philosophical Transactions of the Royal Society B: Biological Sciences 373, 20170434.
137. Johnsson M, Gering E, Willis P, Lopez S, Van Dorp L, Hellenthal G, Henriksen R, Friberg U, Wright D. Feralisation targets different genomic loci to domestication in the chicken. Nature Communications 7, 12950.
138. Kaczensky P, Ganbataar O, Altansukh N, Enkhsaikhan N, Stauffer C, Walzer C. The danger of having all your eggs in one basket – winter crash of the re‐introduced Przewalski's horses in the Mongolian Gobi. PLoS One 6, e28057.
139. Kalbfleisch TS, Rice ES, DePriest MS Jr, Walenz BP, Hestand MS, Vermeesch JR, O Connell BL, Fiddes IT, Vershinina AO, Saremi NF, Petersen JL, Finno CJ, Bellone RR, McCue ME, Brooks SA. Improved reference genome for the domestic horse increases assembly contiguity and composition. Communications Biology 1, 197.
140. Kaseda Y, Khalil AM, Ogawa H. Harem stability and reproductive success of Misaki feral mares. Equine Veterinary Journal 27, 368–372.
141. Kauffmann S, Cline C. The Essential Hoof Book: The Complete Modern Guide to Horse Feet‐Anatomy, Care and Health, Disease Diagnosis and Treatment. Trafalgar Square Books Pomfret, Vermont, 2018.
142. Kay CE. Are ecosystems structured from the top‐down or bottom‐up: a new look at an old debate. Wildlife Society Bulletin 26, 484–498.
143. Kelekna P. The Horse in Human History. Cambridge University Press Cambridge, 2009.
144. Keulartz J. The ethics of wildlife conservation. [Invited lecture] Symposium: Managing Populations of Free‐ranging Herbivores. Utrecht, Netherlands, 2011 .
145. Khan A, Patel K, Bhattacharjee S, Sharma S, Chugani AN, Sivaraman K, Hosawad V, Sahu YK, Reddy GV, Ramakrishnan U. Are shed hair genomes the most effective noninvasive resource for estimating relationships in the wild?. Ecology and Evolution 10, 4583–4594.
146. King SRB, Schoenecker KA, Cole MJ. Effect of adult male sterilization on the behavior and social associations of a feral polygynous ungulate: the horse. Applied Animal Behaviour Science 249, 105598.
147. Klassen L, Rasmussen U, Kveiborg J, Richards M, Orlando L, Svenning J‐C, Ritchie K, Andreasen MH, Philippsen B, Iversen R, Johannsen NN. Ginnerup revisited. New excavations at a key neolithic site on Djursland, Denmark. Journal of Neolithic Archaeology 25, 35–65.
148. Kohl PA, Warner SE. Public communication practices and beliefs among conservation scientists and practitioners. Journal of Fish and Wildlife Management 13, 262–271.
149. Köhler M, Hiller G, Tischew S. Year‐round horse grazing supports typical vascular plant species, orchids and rare bird communities in a dry calcareous grassland. Agriculture, Ecosystems & Environment 234, 48–57.
150. Krueger K. Social ecology of horses. In Ecology of Social Evolution. Springer Berlin, Heidelberg, 2008 .
151. Krüger K, Flauger B. Social feeding decisions in horses (Equus caballus). Behavioural Processes 78, 76–83.
152. Kugler W, Broxham E. The Ecological Value of Feral Livestock Populations in Europe: Overview, Situation and Development of a Network for Management of Wild and Semi‐Wild Livestock Populations, Final Report. SAVE Foundation St. Gallen, 2014.
153. Kveiborg J, Uzunidis A, Klassen L, Klimscha F, Johannsen NN, Rasmussen UL, Richards MP, Svenning J‐C. Paleodiet reconstructions and human utilization of middle Holocene Equus ferus in Northwest Europe. Palaeogeography, Palaeoclimatology, Palaeoecology 649, 112334.
154. Lacy RC. Achieving true sustainability of zoo populations. Zoo Biology 32(1), 19–26.
155. Lagos L, Bárcena F. Spatial variability in wolf diet and prey selection in Galicia (NW Spain). Mammal Research 63, 125–139.
156. Lamoot I, Meert C, Hoffmann M. Habitat use of ponies and cattle foraging together in a coastal dune area. Biological Conservation 122, 523–536.
157. Laseca N, Anaya G, Peña Z, Pirosanto Y, Molina A, Demyda Peyrás S. Impaired reproductive function in equines: from genetics to genomics. Animals 11, 393.
158. Le Busque B, Dorrian J, Litchfield C. The impact of news media portrayals of sharks on public perception of risk and support for shark conservation. Marine Policy 124, 104341.
159. le Roux E, Marneweck DG, Clinning G, Druce DJ, Kerley GIH, Cromsigt JPGM. Top–down limits on prey populations may be more severe in larger prey species, despite having fewer predators. Ecography 42, 1115–1123.
160. Lema JF, Ribeiro S, Palacios V. Observations of wolves hunting free‐ranging horses in NW Iberia. Carnivore Damage Prevention (CDP) News 24, 1–10.
161. Lepetz S, Debue K, Batsukh D. To accompany and honour the deceased: the horses from the graves of the Pazyryk culture. In Masters of the Steppe: The Impact of the Scythians and Later Nomad Societies of Eurasia. Archaeopress Oxford, 2020.
162. Librado P, Gamba C, Gaunitz C, Der Sarkissian C, Pruvost M, Albrechtsen A, Fages A, Khan N, Schubert M, Jagannathan V, Serres‐Armero A, Kuderna LFK, Povolotskaya IS, Seguin‐Orlando A, Lepetz S. Ancient genomic changes associated with domestication of the horse. Science 356, 442–445.
163. Librado P, Khan N, Fages A, Kusliy MA, Suchan T, Tonasso‐Calvière L, Schiavinato S, Alioglu D, Fromentier A, Perdereau A, Aury J‐M, Gaunitz C, Chauvey L, Seguin‐Orlando A, Der Sarkissian C. The origins and spread of domestic horses from the Western Eurasian steppes. Nature 598, 634–640.
164. Librado P, Tressières G, Chauvey L, Fages A, Khan N, Schiavinato S, Calvière‐Tonasso L, Kusliy MA, Gaunitz C, Liu X, Wagner S, Der Sarkissian C, Seguin‐Orlando A, Perdereau A, Aury J‐M. Widespread horse‐based mobility arose around 2200 BCE in Eurasia. Nature 631, 819–824.
165. Linderholm A, Larson G. The role of humans in facilitating and sustaining coat colour variation in domestic animals. Seminars in Cell & Developmental Biology 24, 587–593.
166. Ling B, Raynor EJ, Joern A, Goodin DG. Dynamic plant–herbivore interactions between bison space use and vegetation heterogeneity in a tallgrass prairie. Remote Sensing 15, 5269.
167. Linklater WL, Cameron EZ, Minot EO, Stafford KJ. Stallion harassment and the mating system of horses. Animal Behaviour 58, 295–306.
168. Linnartz L, Meissner R, Lemoine R. Rewilding Horses in Europe: Background and Guidelines. Rewilding Europe Nijmegen, 2023.
169. Linnell JDC, Kaczensky P, Lescureaux N. Human dimensions of wild equid management: exploring the meanings of “wild”. In Wild Equids: Ecology, Management, and Conservation. Johns Hopkins University Press Baltimore, 2016.
170. Liu G, Shafer ABA, Zimmermann W, Hu D, Wang W, Chu H, Cao J, Zhao C. Evaluating the reintroduction project of Przewalski's horse in China using genetic and pedigree data. Biological Conservation 171, 288–298.
171. Lou Hendricks B. International Encyclopedia of Horse Breeds. University of Oklahoma Press Norman, 1995.
172. Lovász L, Fages A, Amrhein V. Konik, Tarpan, European wild horse: an origin story with conservation implications. Global Ecology and Conservation 32, e01911.
173. Lovász L, Korner‐Nievergelt F, Amrhein V. Natural grazing by horses and cattle promotes bird diversity in a restored European alluvial grassland. PeerJ 12, e17777.
174. Lundmark F, Berg C, Wahlberg B, Röcklinsberg H. ‘One animal is no animal’ – consequences of measuring animal welfare at herd level. In Know your Food. Wageningen Academic Publishers Wageningen, 2015.
175. Luthersson N, Nielsen KH, Harris P, Parkin TDH. Risk factors associated with equine gastric ulceration syndrome (EGUS) in 201 horses in Denmark. Equine Veterinary Journal 41, 625–630.
176. Luthersson N, Ýr Þorgrímsdóttir Ú, Harris PA, Parkins T, Bennet ED. Effect of moving from being extensively managed out in pasture into training on the incidence of equine gastric ulcer syndrome in Icelandic horses. Journal of the American Veterinary Medical Association 260, 102110.
177. Mackowski M, Mucha S, Cholewinski G, Cieslak J. Genetic diversity in Hucul and polish primitive horse breeds. Archives Animal Breeding 58, 23–31.
178. Madosky JM, Rubenstein DI, Howard JJ, Stuska S. The effects of immunocontraception on harem fidelity in a feral horse (Equus caballus) population. Applied Animal Behaviour Science 128, 50–56.
179. Malacarne BD, Martins RR, Paz CFR, Alves JVA, Dias LA, Cavalcante MA, Santos AM, Silva AGM, Leise BS, Carvalho AM, Faleiros RR. Histological comparison of the lamellar tissue of Iberian origin breed horses created in semi‐feral conditions or in an intensive system. PLoS One 18, e0286536.
180. Margles SW, Peterson RB, Ervin J, Kaplin BA. Conservation without borders: building communication and action across disciplinary boundaries for effective conservation. Environmental Management 45, 1–4.
181. Matzek V, Covino J, Funk JL, Saunders M. Closing the knowing–doing gap in invasive plant management: accessibility and interdisciplinarity of scientific research. Conservation Letters 7, 208–215.
182. McCue ME, Bannasch DL, Petersen JL, Gurr J, Bailey E, Binns MM, Distl O, Guérin G, Hasegawa T, Hill EW, Leeb T, Lindgren G, Penedo MCT, Røed KH, Ryder OA. A high density SNP array for the domestic horse and extant Perissodactyla: utility for association mapping, genetic diversity, and phylogeny studies. PLoS Genetics 8, e1002451.
183. McLeod SR. Is the concept of carrying capacity useful in variable environments?. Oikos 79, 529–542.
184. Mellish MA, Lucas ZN, Puchalski SM, Kusch TA. An estimation of lameness in sable Island horses using radiographic evaluation of the distal phalanx and hoof capsule. Journal of Applied Animal Welfare Science 26, 184–194.
185. Meserve PL, Kelt DA, Milstead WB, Gutiérrez JR. Thirteen years of shifting top‐down and bottom‐up control. Bioscience 53, 633–646.
186. Metcalf JL, Song SJ, Morton JT, Weiss S, Seguin‐Orlando A, Joly F, Feh C, Taberlet P, Coissac E, Amir A, Willerslev E, Knight R, McKenzie V, Orlando L. Evaluating the impact of domestication and captivity on the horse gut microbiome. Scientific Reports 7, 15497.
187. Michelot M. Origine, statut et perspectives. Revue Scientifique Bourgogne‐Nature 21, 348–357.
188. Mihlbachler MC, Rivals F, Solounias N, Semprebon GM. Dietary change and evolution of horses in North America. Science 331, 1178–1181.
189. Moinardeau C, Mesleard F, Ramone H, Dutoit T. Grazing in temporary paddocks with hardy breed horses (Konik polski) improved species‐rich grasslands restoration in artificial embankments of the Rhône river (Southern France). Global Ecology and Conservation 31, 1874.
190. Monthoux C, de Brot S, Jackson M, Bleul U, Walter J. Skin malformations in a neonatal foal tested homozygous positive for warmblood fragile foal syndrome. BMC Veterinary Research 11, 12.
191. Morgan R, Keen J, McGowan C. Equine metabolic syndrome. The Veterinary Record 177, 173–179.
192. Murchie TJ, Monteath AJ, Mahony ME, Long GS, Cocker S, Sadoway T, Karpinski E, Zazula G, MacPhee RDE, Froese D, Poinar HN. Collapse of the mammoth‐steppe in central Yukon as revealed by ancient environmental DNA. Nature Communications 12, 7120.
193. Murray MJ, Eichorn ES. Effects of intermittent feed deprivation, intermittent feed deprivation with ranitidine administration, and stall confinement with ad libitum access to hay on gastric ulceration in horses. American Journal of Veterinary Research 57, 1599–1603.
194. Naundrup PJ, Svenning J‐C. A geographic assessment of the global scope for rewilding with wild‐living horses (Equus ferus). PLoS One 10, e0132359.
195. Notzke C. An exploration into political ecology and nonhuman agency: the case of the wild horse in Western Canada. The Canadian Geographer 57, 389–412.
196. Notzke C. Wild horse‐based tourism as wildlife tourism: the wild horse as the other. Current Issues in Tourism 19, 1235–1259.
197. Nuñez CMV, Adelman JS, Carr HA, Alvarez CM, Rubenstein DI. Lingering effects of contraception management on feral mare (Equus caballus) fertility and social behavior. Conservation Physiology 5, cox018.
198. Nuñez CMV, Scorolli A, Lagos L, Berman D, Kane AJ. Management of free‐roaming horses. In Wild Equids: Ecology, Management, and Conservation. Johns Hopkins University Press Baltimore, 2016.
199. Orlando L, Ginolhac A, Zhang G, Froese D, Albrechtsen A, Stiller M, Schubert M, Cappellini E, Petersen B, Moltke I, Johnson PLF, Fumagalli M, Vilstrup JT, Raghavan M, Korneliussen T. Recalibrating Equus evolution using the genome sequence of an early Middle Pleistocene horse. Nature 499, 74–78.
200. Orlando L, Librado P. Origin and evolution of deleterious mutations in horses. Genes 10, 649.
201. Outram AK. Horse domestication as a multi‐centered, multi‐stage process: Botai and the role of specialized Eneolithic horse pastoralism in the development of human‐equine relationships. Frontiers in Environmental Archaeology 2, 1134068.
202. Outram AK, Stear NA, Bendrey R, Olsen S, Kasparov A, Zaibert V, Thorpe N, Evershed RP. The earliest horse harnessing and milking. Science 323, 1332–1335.
203. Pausas JG, Bond WJ. Humboldt and the reinvention of nature. Journal of Ecology 107, 1031–1037.
204. Pearce EA, Mazier F, Normand S, Fyfe R, Andrieu V, Bakels C, Balwierz Z, Bińka K, Boreham S, Borisova OK, Brostrom A. Substantial light woodland and open vegetation characterized the temperate forest biome before Homo sapiens. Science Advances 9, eadi9135.
205. Perino A, Pereira HM, Navarro LM, Fernández N, Bullock JM, Ceaușu S, Cortés‐Avizanda A, van Klink R, Kuemmerle T, Lomba A, Pe'er G, Plieninger T, Rey Benayas JM, Sandom CJ, Svenning J‐C. Rewilding complex ecosystems. Science 364, eaav5570.
206. Petersen SL, Scasta JD, Schoenecker KA, Hennig JD. Feral equids. In Rangeland Wildlife Ecology and Conservation. Springer Nature Switzerland AG Cham, 2023.
207. Peterson J. Ungulate/Vegetation Dynamics in the Pryor Mountain Wild Horse Range. Colorado State University Fort Collins, CO, 1999.
208. Pettorelli N, Barlow J, Stephens PA, Durant SM, Connor B, Schulte to Bühne H, Sandom CJ, Wentworth J, du Toit JT. Making rewilding fit for policy. The Journal of Applied Ecology 55, 1114–1125.
209. Pruvost M, Bellone R, Benecke N, Sandoval‐Castellanos E, Cieslak M, Kuznetsova T, Morales‐Muñiz A, O'Connor T, Reissmann M, Hofreiter M, Ludwig A. Genotypes of predomestic horses match phenotypes painted in Paleolithic works of cave art. Proceedings of the National Academy of Sciences of the United States of America 108, 18626–18630.
210. Ramp D, Bekoff M. Compassion as a practical and evolved ethic for conservation. In Animal Ethics in the Age of Humans: Blurring Boundaries in Human‐Animal Relationships. Springer Nature Switzerland AG Cham, 2016.
211. Ransom JI, Hobbs NT, Bruemmer J. Contraception can lead to trophic asynchrony between birth pulse and resources. PLoS One 8, e54972.
212. Ransom JI, Kaczensky P. Wild Equids: Ecology, Management, and Conservation. Johns Hopkins University Press Baltimore, 2016.
213. Ransom JI, Lagos LA, Hrabar HA, Nowzari HA, Usukhjargal D, Spasskaya N. Wild and feral equid population dynamics. In Wild Equids: Ecology, Management, and Conservation. Johns Hopkins University Press Baltimore, 2016.
214. Ransom JI, Powers JG, Thompson Hobbs N, Baker DL. Ecological feedbacks can reduce population‐level efficacy of wildlife fertility control. Journal of Applied Ecology 51, 259–269.
215. Raudsepp T, Finno CJ, Bellone RR, Petersen JL. Ten years of the horse reference genome: insights into equine biology, domestication and population dynamics in the post‐genome era. Animal Genetics 50, 569–597.
216. Richardson RC. Self‐Healing for the Horse's Foot. A Site for Barefoot and Shod Horses, Rockfoot Retrieved 10 October, 2023 from https://www.rockfoot.com/self_healing.html. .
217. Rikoon JS. Wild horses and the political ecology of nature restoration in the Missouri Ozarks. Geoforum 37, 200–211.
218. Ripple WJ, Beschta RL. Wolves and the ecology of fear: can predation risk structure ecosystems?. Bioscience 54, 755–766.
219. Robinson IH. The human‐horse relationship: how much do we know?. Equine Veterinary Journal 31, 42–45.
220. Rogers GM. Kaimanawa feral horses and their environmental impacts. New Zealand Journal of Ecology 15, 49–64.
221. Römbke J, Coors A, Fernández ÁA, Förster B, Fernández C, Jensen J, Lumaret JP, Cots MÁP, Liebig M. Effects of the parasiticide ivermectin on the structure and function of dung and soil invertebrate communities in the field (Madrid, Spain). Applied Soil Ecology 45, 284–292.
222. Rook L, Bernor RL, Avilla LS, Cirilli O, Flynn L, Jukar A, Sanders W, Scott E, Wang X. Mammal biochronology (land mammal ages) around the world from late Miocene to middle Pleistocene and major events in horse evolutionary history. Frontiers in Ecology and Evolution 7, 278.
223. Rosa LP, Mallicote MF, Long MT, Brooks SA. Metabogenomics reveals four candidate regions involved in the pathophysiology of Equine Metabolic Syndrome. Molecular and Cellular Probes 53, 101620.
224. Rudolph JA, Spier SJ, Byrns G, Rojas CV, Bernoco D, Hoffman EP. Periodic paralysis in quarter horses: a sodium channel mutation disseminated by selective breeding. Nature Genetics 2, 144–147.
225. Rutberg AT, Turner JW Jr, Herman. Fertility control and the welfare of free‐roaming horses and burros on U.S. public lands: the need for an ethical framing. Animals 12, 2656.
226. Saavedra D, Fernández N, Svenning J‐C. Addressing challenges for large‐scale trophic rewilding. Journal for Nature Conservation 73, 126382.
227. Sairanen J, Nivola K, Katila T, Virtala AM, Ojala M. Effects of inbreeding and other genetic components on equine fertility. Animal 3, 1662–1672.
228. Sala N, Alcaraz‐Castaño M, Arriolabengoa M, Martínez‐Pillado V, Pantoja‐Pérez A, Rodríguez‐Hidalgo A, Téllez E, Cubas M, Castillo S, Arnold LJ, Demuro M. Nobody's land? The oldest evidence of early Upper Paleolithic settlements in inland Iberia. Science Advances 10, 3807.
229. Sandom CJ, Ejrnæs R, Hansen MDD, Svenning J‐C. High herbivore density associated with vegetation diversity in interglacial ecosystems. Proceedings of the National Academy of Sciences of the United States of America 111, 4162–4167.
230. Särkijärvi S, Reilas T, Saastamoinen M, Elo K, Jaakkola S, Kokkonen T. Effect of cultivated or semi‐natural pasture on changes in live weight, body condition score, body measurements and fat thickness in grazing Finnhorse mares. In Forages and Grazing in Horse Nutrition. Wageningen Academic Publishers Wageningen, 2012.
231. Sas‐Jaworsky A, Scasta JD. Wyoming's wild horse ranch: history and description of a socio‐ecological experiment. Human–Wildlife Interactions 16, 13.
232. Sauvet G. The hierarchy of animals in the Paleolithic iconography. Journal of Archaeological Science: Reports 28, 102025.
233. Scasta JD. Why are humans so emotional about feral horses? A spatiotemporal review of the psycho‐ecological evidence with global implications. Geoforum 103, 171–175.
234. Scasta JD, Adams M, Gibbs R, Fleury B. Free‐ranging horse management in Australia, New Zealand and the United States: socio‐ecological dimensions of a protracted environmental conflict. Rangeland Journal 42, 27–43.
235. Scasta JD, Beck JL, Angwin CJ. Meta‐analysis of diet composition and potential conflict of wild horses with livestock and wild ungulates on western rangelands of North America. Rangeland Ecology & Management 69, 310–318.
236. Scasta JD, Thacker E, Hennig JD, Hoopes K. Dehydration and mortality of feral horses and burros. Human‐Wildlife Interactions 16, 251–261.
237. Schaefer RJ, McCue ME. Equine genotyping arrays. The Veterinary Clinics of North America: Equine Practice 36, 183–193.
238. Schaefer RJ, Schubert M, Bailey E, Bannasch DL, Barrey E, Bar‐Gal GK, Brem G, Brooks SA, Distl O, Fries R, Finno CJ, Gerber V, Haase B, Jagannathan V, Kalbfleisch T. Developing a 670k genotyping array to tag ~2M SNPs across 24 horse breeds. BMC Genomics 18, 565.
239. Schmitz OJ, Sylvén M, Atwood TB, Bakker ES, Berzaghi F, Brodie JF, Cromsigt JP, Davies AB, Leroux SJ, Schepers FJ, Smith FA. Trophic rewilding can expand natural climate solutions. Nature Climate Change 13, 324–333.
240. Schweizer RM, Saarman N, Ramstad KM, Forester BR, Kelley JL, Hand BK, Malison RL, Ackiss AS, Watsa M, Nelson TC, Beja‐Pereira A, Waples RS, Funk WC, Luikart G. Big data in conservation genomics: boosting skills, hedging bets, and staying current in the field. The Journal of Heredity 112, 313–327.
241. Senn J. Large herbivores. In Disturbance Ecology. Springer Nature Switzerland AG Cham, 2022.
242. Shafer ABA, Wolf JBW, Alves PC, Bergström L, Bruford MW, Brännström I, Colling G, Dalén L, De Meester L, Ekblom R, Fawcett KD, Fior S, Hajibabaei M, Hill JA, Hoezel AR. Genomics and the challenging translation into conservation practice. Trends in Ecology & Evolution 30, 78–87.
243. Sigurjónsdóttir H, van Dierendonck M, Snorrason S, Thórhallsdóttir A. Social relationships in a group of horses without a mature stallion. Behaviour 140, 783–804.
244. Simonin D, Gavinelli A. The European Union legislation on animal welfare: state of play, enforcement and future activities. In Animal Welfare: From Science to Law. La Fondation Droit Animal, Éthique & Sciences Paris, 2019.
245. Skogland T. What are the effects of predators on large ungulate populations?. Oikos 61, 401–411.
246. Soulé ME. What is conservation biology? A new synthetic discipline addresses the dynamics and problems of perturbed species, communities, and ecosystems. Bioscience 35, 727–734.
247. Steelman SM, Chowdhary BP, Dowd S, Suchodolski J, Janečka JE. Pyrosequencing of 16S rRNA genes in fecal samples reveals high diversity of hindgut microflora in horses and potential links to chronic laminitis. BMC Veterinary Research 8, 231.
248. Stevens EF. Contests between bands of feral horses for access to fresh water: the resident wins. Animal Behaviour 36, 1851–1853.
249. Suchorski‐Tremblay A, Kok R, Thomason R. Modelling horse hoof cracking with artificial neural networks. Canadian Biosystems Engineering 43, 7.15–7.22.
250. Supple MA, Shapiro B. Conservation of biodiversity in the genomics era. Genome Biology 19, 131.
251. Sutherland WJ, Spiegelhalter D, Burgman MA. Policy: twenty tips for interpreting scientific claims. Nature 503, 335–337.
252. Svenning JC, Buitenwerf R, Le Roux E. Trophic rewilding as a restoration approach under emerging novel biosphere conditions. Current Biology 34, 435–451.
253. Svenning J‐C, Pedersen PBM, Donlan CJ, Ejrnæs R, Faurby S, Galetti M, Hansen DM, Sandel B, Sandom CJ, Terborgh JW, Vera FWM. Science for a wilder Anthropocene: synthesis and future directions for trophic rewilding research. Proceedings of the National Academy of Sciences of the United States of America 113, 898–906.
254. Swegen A, Aitken RJ. Prospects for immunocontraception in feral horse population control: exploring novel targets for an equine fertility vaccine. Reproduction, Fertility, and Development 28, 853–863.
255. Szmatoła T, Gurgul A, Jasielczuk I, Oclon E, Ropka‐Molik K, Stefaniuk‐Szmukier M, Polak G, Tomczyk‐Wrona I, Bugno‐Poniewierska M. Assessment and distribution of runs of homozygosity in horse breeds representing different utility types. Animals 12, 3293.
256. Tanner E, White A, Lurz PWW, Gortázar C, Díez‐Delgado I, Boots M. The critical role of infectious disease in compensatory population growth in response to culling. The American Naturalist 194, E1–E12.
257. Taylor HR, Dussex N, van Heezik Y. Bridging the conservation genetics gap by identifying barriers to implementation for conservation practitioners. Global Ecology and Conservation 10, 231–242.
258. Taylor RS, Manseau M, Redquest B, Keobouasone S, Gagné P, Martineau C, Wilson PJ. Whole genome sequences from non‐invasively collected caribou faecal samples. Conservation Genetics Resources 14, 53–68.
259. Taylor WTT, Barrón‐Ortiz CI. Rethinking the evidence for early horse domestication at Botai. Scientific Reports 11, 7440.
260. Theunissen B. The Oostvaardersplassen Fiasco. Isis 110, 341–345.
261. Thulin C‐G, Alves PC, Djan M, Fontanesi L, Peacock D. Wild opportunities with dedomestication genetics of rabbits. Restoration Ecology 25, 330–332.
262. Thulin C‐G, Röcklinsberg H. Ethical considerations for wildlife reintroductions and rewilding. Frontiers in Veterinary Science 7, 163.
263. Timmermann A, Damgaard C, Strandberg MT, Svenning J‐C. Pervasive early 21st‐century vegetation changes across Danish semi‐natural ecosystems: more losers than winners and a shift towards competitive, tall‐growing species. Journal of Applied Ecology 52, 21–30.
264. Tree I. Wilding: The Return of Nature to a British Farm. Pan Macmillan UK, 2018.
265. Treiber KH, Boston RC, Kronfeld DS, Staniar WB, Harris PA. Insulin resistance and compensation in Thoroughbred weanlings adapted to high‐glycemic meals. Journal of Animal Science 83, 2357–2364.
266. Treiber KH, Kronfeld DS, Hess TM, Byrd BM, Splan RK, Staniar WB. Evaluation of genetic and metabolic predispositions and nutritional risk factors for pasture‐associated laminitis in ponies. Journal of the American Veterinary Medical Association 228, 1538–1545.
267. Trouwborst A, Svenning J‐C. Megafauna restoration as a legal obligation: international biodiversity law and the rehabilitation of large mammals in Europe. Review of European, Comparative & International Environmental Law 31, 182–198.
268. Turghan MA, Jiang Z, Niu Z. An update on status and conservation of the Przewalski's horse (Equus ferus przewalskii): captive breeding and reintroduction projects. Animals 12, 3158.
269. Tydén E, Jansson A, Ringmark S. Parasites in horses kept in a 2.5 year‐round grazing system in nordic conditions without supplementary feeding. Animals 9, 1156.
270. Tyler SJ. The behaviour and social organization of the new forest ponies. Animal Behaviour Monographs 5, 87–196.
271. Ubilla M, Rinderknecht A, Corona A, Perea D. Mammals in last 30 to 7 ka interval (late Pleistocene–early Holocene) in Southern Uruguay (Santa Lucía River Basin): last occurrences, climate, and biogeography. Journal of Mammalian Evolution 25, 291–300.
272. Urban B, Thieme H, Elsner H. Biostratigraphische, quartärgeologische und urgeschichtliche Befunde aus dem Tagebau. Zeitschrift der Deutschen Morgenlandischen Gesellschaft 139, 123–154.
273. Urban L, Miller AK, Eason D, Vercoe D, Shaffer M, Wilkinson SP, Jeunen G‐J, Gemmell NJ, Digby A. Non‐invasive real‐time genomic monitoring of the critically endangered kākāpō. eLife 12, RP84553.
274. van den Boom R. Equine gastric ulcer syndrome in adult horses. Veterinary Journal 283‐284, 105830.
275. van Dierendonck MC. The Importance of Social Relationships in Horses. Utrecht University Utrecht, 2006.
276. van Dierendonck MC, Spruijt BM. Coping in groups of domestic horses – review from a social and neurobiological perspective. Applied Animal Behaviour Science 138(3–4), 194–202.
277. van Eeden LM, Newsome TM, Crowther MS, Dickman CR, Bruskotter J. Social identity shapes support for management of wildlife and pests. Biological Conservation 231, 167–173.
278. van Klink R, van der Plas F, van Noordwijk CGET, Wallis DeVries MF, Olff H. Effects of large herbivores on grassland arthropod diversity. Biological Reviews of the Cambridge Philosophical Society 90, 347–366.
279. Veit W, Browning H. Extending animal welfare science to include wild animals. Animal Sentience 1, 20.
280. Verga M, Michelazzi M. Companion animal welfare and possible implications on the human–pet relationship. Italian Journal of Animal Science 8, 231–240.
281. Vermeulen R. Natural Grazing: Practices in the Rewilding of Cattle and Horses. Rewilding Europe Nijmegen, 2015.
282. Vershinina AO, Heintzman PD, Froese DG, Zazula G, Cassatt‐Johnstone M, Dalén L, Der Sarkissian C, Dunn SG, Ermini L, Gamba C, Groves P, Kapp JD, Mann DH, Seguin‐Orlando A, Southon J. Ancient horse genomes reveal the timing and extent of dispersals across the Bering Land Bridge. Molecular Ecology 30, 6144–6161.
283. Vincelette AR, Renders E, Scott KM, Falkingham PL, Janis CM. Hipparion tracks and horses' toes: the evolution of the equid single hoof. Royal Society Open Science 10, 230358.
284. von Thaden A, Nowak C, Tiesmeyer A, Reiners TE, Alves PC, Lyons LA, Mattucci F, Randi E, Cragnolini M, Galián J, Hegyeli Z, Kitchener AC, Lambinet C, Lucas JM, Mölich T. Applying genomic data in wildlife monitoring: development guidelines for genotyping degraded samples with reduced single nucleotide polymorphism panels. Molecular Ecology Resources 20, 662–680.
285. Voogt AM, Ursinus WW, Sijm DTHM, Bongers JH. From the Five Freedoms to a more holistic perspective on animal welfare in the Dutch animals act. Frontiers in Animal Science 4, 1026224.
286. Wallner A, Hunziker M. The wolf controversy ‐ interviews with experts on the social acceptance of the wolf in Switzerland. Forest Snow and Landscape Research 76, 191–212.
287. Wallner B, Palmieri N, Vogl C, Rigler D, Bozlak E, Druml T, Jagannathan V, Leeb T, Fries R, Tetens J, Thaller G, Metzger J, Distl O, Lindgren G, Rubin CJ. Y chromosome uncovers the recent oriental origin of modern stallions. Current Biology 27, 2029–2035.
288. Walsh JC, Dicks LV, Sutherland WJ. The effect of scientific evidence on conservation practitioners' management decisions. Conservation Biology 29, 88–98.
289. Wang X, Pipes L, Trut LN, Herbeck Y, Vladimirova AV, Gulevich RG, Kharlamova AV, Johnson JL, Acland GM, Kukekova AV, Clark AG. Genomic responses to selection for tame/aggressive behaviors in the silver fox (Vulpes vulpes). Proceedings of the National Academy of Sciences of the United States of America 115, 10398–10403.
290. Ward A, Webster M. Sociality: The Behaviour of Group‐Living Animals. Springer Berlin, 2016.
291. Ward S, Sykes BW, Brown H, Bishop A, Penaluna LA. A comparison of the prevalence of gastric ulceration in feral and domesticated horses in the UK. Equine Veterinary Education 27, 655–657.
292. Waring G. Horse Behavior. The Behavioral Traits and Adaptations of Domestic and Wild Horses, Including Ponies. Noyes Publications Park Ridge, 1983.
293. Waters MR, Stafford TW Jr, Kooyman B, Hills LV. Late Pleistocene horse and camel hunting at the southern margin of the ice‐free corridor: reassessing the age of Wally's beach, Canada. Proceedings of the National Academy of Sciences of the United States of America 112, 4263–4267.
294. Webster AJ. Farm animal welfare: the five freedoms and the free market. Veterinary Journal 161, 229–237.
295. Wesche K, Krause B, Culmsee H, Leuschner C. Fifty years of change in central European grassland vegetation: large losses in species richness and animal‐pollinated plants. Biological Conservation 150, 76–85.
296. Wutke S, Sandoval‐Castellanos E, Benecke N, Döhle H‐J, Friederich S, Gonzalez J, Hofreiter M, Lõugas L, Magnell O, Malaspinas A‐S, Morales‐Muñiz A, Orlando L, Reissmann M, Trinks A, Ludwig A. Decline of genetic diversity in ancient domestic stallions in Europe. Science Advances 4, eaap9691.
297. Yarnell K, Hall C, Royle C, Walker SL. Domesticated horses differ in their behavioural and physiological responses to isolated and group housing. Physiology & Behavior 143, 51–57.
298. Zahl‐Thanem A, Burton RJF, Blekesaune A, Haugen MS, Rønningen K. The impact of wolves on psychological distress among farmers in Norway. Journal of Rural Studies 78, 1–11.
299. Zimmermann W. International Przewalski's Horse Studbook (Equus ferus przewalskii). Cologne Zoo Cologne, 2014.

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