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Topic:Arboviruses

Arboviruses are a group of viruses transmitted by arthropods, such as mosquitoes and ticks, that can infect horses, leading to various diseases. These viruses include the West Nile virus, Eastern equine encephalitis virus, and Western equine encephalitis virus, among others. Infection with arboviruses can result in neurological symptoms in horses, including ataxia, fever, and in severe cases, encephalitis. The transmission dynamics, pathogenesis, and clinical outcomes of arboviral infections in horses are subjects of ongoing research. This page compiles peer-reviewed research studies and scholarly articles that explore the epidemiology, clinical manifestations, diagnostic methods, and preventive measures related to arboviral infections in equine populations.
Equine viral encephalitis: prevalence, impact, and management strategies.
Veterinary medicine (Auckland, N.Z.)    August 7, 2019   Volume 10 99-110 doi: 10.2147/VMRR.S168227
Barba M, Fairbanks EL, Daly JM.Members of several different virus families cause equine viral encephalitis, the majority of which are arthropod-borne viruses (arboviruses) with zoonotic potential. The clinical signs caused are rarely pathognomonic; therefore, a clinical diagnosis is usually presumptive according to the geographical region. However, recent decades have seen expansion of the geographical range and emergence in new regions of numerous viral diseases. In this context, this review presents an overview of the prevalence and distribution of the main viral causes of equine encephalitis and discusses their impact an...
Serological evidence of infection with dengue and Zika viruses in horses on French Pacific Islands.
PLoS neglected tropical diseases    February 7, 2019   Volume 13, Issue 2 e0007162 doi: 10.1371/journal.pntd.0007162
Beck C, Leparc-Goffart I, Desoutter D, Debergé E, Bichet H, Lowenski S, Dumarest M, Gonzalez G, Migné C, Vanhomwegen J, Zientara S, Durand B....New Caledonia and French Polynesia are areas in which arboviruses circulate extensively. A large serological survey among horses from New Caledonia and French Polynesia was carried out to investigate the seroprevalence of flaviviruses in the horse population. Here, 293 equine sera samples were screened for flaviviruses using a competitive enzyme-linked immunosorbent assay (cELISA). The positive sera were then confirmed using a flavivirus-specific microsphere immunoassay (MIA) and seroneutralization tests. This serosurvey showed that 16.6% (27/163) and 30.8% (40/130) of horses were positive for...
West Nile virus infection in horses in Saudi Arabia (in 2013-2015).
Zoonoses and public health    November 6, 2018   Volume 66, Issue 2 248-253 doi: 10.1111/zph.12532
Hemida MG, Perera RAPM, Chu DKW, Ko RLW, Alnaeem AA, Peiris M.West Nile virus (WNV) is an important emerging zoonotic arbovirus giving rise to clinical syndromes of varying severity in humans and horses. Culex mosquitoes are the main vector. Although WNV has been reported in many countries in the Middle East and Asia, little is known about its prevalence in equine populations in the Arabian Peninsula. We have carried out a serological study on 200 horses to assess WNV infection in the Eastern and Central regions of Saudi Arabia in 2013-2015. Sera were tested for the presence of WNV antibodies in parallel using a commercial enzyme-linked immunosorbent ass...
West Nile virus associated with equid encephalitis in Brazil, 2018.
Transboundary and emerging diseases    November 2, 2018   Volume 66, Issue 1 445-453 doi: 10.1111/tbed.13043
Silva ASG, Matos ACD, da Cunha MACR, Rehfeld IS, Galinari GCF, Marcelino SAC, Saraiva LHG, Martins NRDS, Maranhão RPA, Lobato ZIP, Pierezan F....Mosquito-borne arboviruses are a major public health concern worldwide and are responsible for emerging and re-emerging diseases. Taken together, the arboviruses have a strong impact on public health and are the most common causes of equine encephalitis. In-depth diagnostic investigation of equine viral encephalitis is of utmost importance for the epidemiological surveillance and control of this disease. Regarding neurological disorders in equids, in April-May 2018, at least 12 cases of equid mortality with acute neurological signs were reported in six farms from Espirito Santo state, Brazil. ...
Population genomics of Culiseta melanura, the principal vector of Eastern equine encephalitis virus in the United States.
PLoS neglected tropical diseases    August 17, 2018   Volume 12, Issue 8 e0006698 doi: 10.1371/journal.pntd.0006698
Soghigian J, Andreadis TG, Molaei G.Eastern Equine Encephalitis (EEE) (Togaviridae, Alphavirus) is a highly pathogenic mosquito-borne arbovirus that circulates in an enzootic cycle involving Culiseta melanura mosquitoes and wild Passeriformes birds in freshwater swamp habitats. Recently, the northeastern United States has experienced an intensification of virus activity with increased human involvement and northward expansion into new regions. In addition to its principal role in enzootic transmission of EEE virus among avian hosts, recent studies on the blood-feeding behavior of Cs. melanura throughout its geographic range sugg...
Seroprevalence and associated risk factors of mosquito-borne alphaviruses in horses in northern Queensland.
Australian veterinary journal    June 27, 2018   Volume 96, Issue 7 243-251 doi: 10.1111/avj.12711
Gummow B, Tan R, Joice RK, Burgess G, Picard J.To investigate the seroprevalence and associated risk factors of alphaviruses (Ross River virus (RRV), Barmah Forest virus (BFV) and Whataroa virus (WHAV)) in northern Queensland horses. Methods: A cross-sectional study of alphavirus antibodies in horses (n = 287) from 147 properties in northern Queensland from September 2013 to June 2014 was conducted. Owners of sampled horses were interviewed on potential risk factors. Data were analysed for associations using multivariable logistic regression. Results: Antibody titres for RRV were demonstrated in samples from 134 properties (91%; 95% co...
Survey of UK horse owners’ knowledge of equine arboviruses and disease vectors.
The Veterinary record    May 15, 2018   Volume 183, Issue 5 159 doi: 10.1136/vr.104521
Chapman GE, Baylis M, Archer DC.Increased globalisation and climate change have led to concern about the increasing risk of arthropod-borne virus (arbovirus) outbreaks globally. An outbreak of equine arboviral disease in northern Europe could impact significantly on equine welfare, and result in economic losses. Early identification of arboviral disease by horse owners may help limit disease spread. In order to determine what horse owners understand about arboviral diseases of horses and their vectors, the authors undertook an open, cross-sectional online survey of UK horse owners. The questionnaire was distributed using soc...
The challenges posed by equine arboviruses.
Equine veterinary journal    April 14, 2018   Volume 50, Issue 4 436-445 doi: 10.1111/evj.12829
Chapman GE, Baylis M, Archer D, Daly JM.Equine populations worldwide are at increasing risk of infection by viruses transmitted by biting arthropods, including mosquitoes, biting midges (Culicoides), sandflies and ticks. These include the flaviviruses (Japanese encephalitis, West Nile and Murray Valley encephalitis), alphaviruses (eastern, western and Venezuelan encephalitis) and the orbiviruses (African horse sickness and equine encephalosis). This review provides an overview of the challenges faced in the surveillance, prevention and control of the major equine arboviruses, particularly in the context of these viruses emerging in ...
The Immune Responses of the Animal Hosts of West Nile Virus: A Comparison of Insects, Birds, and Mammals.
Frontiers in cellular and infection microbiology    April 3, 2018   Volume 8 96 doi: 10.3389/fcimb.2018.00096
Ahlers LRH, Goodman AG.Vector-borne diseases, including arboviruses, pose a serious threat to public health worldwide. Arboviruses of the flavivirus genus, such as Zika virus (ZIKV), dengue virus, yellow fever virus (YFV), and West Nile virus (WNV), are transmitted to humans from insect vectors and can cause serious disease. In 2017, over 2,000 reported cases of WNV virus infection occurred in the United States, with two-thirds of cases classified as neuroinvasive. WNV transmission cycles through two different animal populations: birds and mosquitoes. Mammals, particularly humans and horses, can become infected thro...
Alphaviruses: Serological Evidence of Human Infection in Paraguay (2012-2013).
Vector borne and zoonotic diseases (Larchmont, N.Y.)    March 13, 2018   Volume 18, Issue 5 266-272 doi: 10.1089/vbz.2017.2178
Alphaviruses can produce febrile illness and encephalitis in dead-end hosts such as horses and humans. Within this genus, the Venezuelan Equine Encephalitis virus (VEEV) complex includes pathogenic epizootic subtypes and enzootic subtypes that are not pathogenic in horses (except subtype IE, Mexican strains), although they can cause febrile symptoms in humans. The Rio Negro virus (RNV-VEEV subtype VI) circulates in Argentina, where it was associated with undifferentiated febrile illness. Mayaro (MAYV) and Una (UNAV) viruses belong to a different group, the Semliki Forest virus complex, with co...
Abundance and species composition of Culicoides spp. biting midges near cattle and horse in South-Eastern Poland.
Acta parasitologica    October 17, 2017   Volume 62, Issue 4 739-747 doi: 10.1515/ap-2017-0089
Larska M, Grochowska M, Lechowski L, Żmudziński JF.The aim of the study was to estimate and compare the distribution of Culicoides biting midges species at farms with different main hosts - cattle and horse. Culicoides spp. are known vectors of arboviruses including African horse sickness virus (AHSV), bluetongue virus (BTV) and Schmallenberg virus (SBV). The latter two have been already reported in Polish ruminants recently, while AHSV remains absent, however the risk of its emergence has been increasing in the recent years. In order to establish the activity of potential AHSV vector at vicinity of horses, an OVI midge trap has been placed at...
Emerging arboviruses in Quebec, Canada: assessing public health risk by serology in humans, horses and pet dogs.
Epidemiology and infection    September 28, 2017   Volume 145, Issue 14 2940-2948 doi: 10.1017/S0950268817002205
Rocheleau JP, Michel P, Lindsay LR, Drebot M, Dibernardo A, Ogden NH, Fortin A, Arsenault J.Periodic outbreaks of West Nile virus (WNV), Eastern equine encephalitis virus (EEEV) and to a lesser extent, California serogroup viruses (CSGV), have been reported in parts of Canada in the last decade. This study was designed to provide a broad assessment of arboviral activity in Quebec, Canada, by conducting serological surveys for these arboviruses in 196 horses, 1442 dogs and 485 humans. Sera were screened by a competitive enzyme linked immunosorbent assay and positive samples confirmed by plaque reduction neutralisation tests. The percentage of seropositive samples was 83·7%, 16·5%, 7...
Evolution and spread of Venezuelan equine encephalitis complex alphavirus in the Americas.
PLoS neglected tropical diseases    August 3, 2017   Volume 11, Issue 8 e0005693 doi: 10.1371/journal.pntd.0005693
Forrester NL, Wertheim JO, Dugan VG, Auguste AJ, Lin D, Adams AP, Chen R, Gorchakov R, Leal G, Estrada-Franco JG, Pandya J, Halpin RA, Hari K, Jain R....Venezuelan equine encephalitis (VEE) complex alphaviruses are important re-emerging arboviruses that cause life-threatening disease in equids during epizootics as well as spillover human infections. We conducted a comprehensive analysis of VEE complex alphaviruses by sequencing the genomes of 94 strains and performing phylogenetic analyses of 130 isolates using complete open reading frames for the nonstructural and structural polyproteins. Our analyses confirmed purifying selection as a major mechanism influencing the evolution of these viruses as well as a confounding factor in molecular cloc...
Characterization of Fitzroy River Virus and Serologic Evidence of Human and Animal Infection.
Emerging infectious diseases    July 21, 2017   Volume 23, Issue 8 1289-1299 doi: 10.3201/eid2308.161440
Johansen CA, Williams SH, Melville LF, Nicholson J, Hall RA, Bielefeldt-Ohmann H, Prow NA, Chidlow GR, Wong S, Sinha R, Williams DT, Lipkin WI....In northern Western Australia in 2011 and 2012, surveillance detected a novel arbovirus in mosquitoes. Genetic and phenotypic analyses confirmed that the new flavivirus, named Fitzroy River virus, is related to Sepik virus and Wesselsbron virus, in the yellow fever virus group. Most (81%) isolates came from Aedes normanensis mosquitoes, providing circumstantial evidence of the probable vector. In cell culture, Fitzroy River virus replicated in mosquito (C6/36), mammalian (Vero, PSEK, and BSR), and avian (DF-1) cells. It also infected intraperitoneally inoculated weanling mice and caused mild c...
Circulation of Zoonotic Arboviruses in Equine Populations of Mallorca Island (Spain).
Vector borne and zoonotic diseases (Larchmont, N.Y.)    March 27, 2017   Volume 17, Issue 5 340-346 doi: 10.1089/vbz.2016.2042
Vanhomwegen J, Beck C, Desprès P, Figuerola A, García R, Lecollinet S, López-Roig M, Manuguerra JC, Serra-Cobo J.The presence of major arbovirus vector species, climate change that promotes the expansion and increase of their populations, and potential animal reservoirs mean that vector-borne diseases represent a significant health risk for Mallorca's inhabitants. Microbiological monitoring of circulating arboviruses, particularly flaviviruses causing encephalitis, was initiated using domestic horses from localities near wetlands as "sentinel" hosts. A total of 291 blood samples were taken from 172 horses between 2011 and 2012, using paired samples to highlight seroconversion events. A multiplex immunoas...
African Horse Sickness Virus: History, Transmission, and Current Status.
Annual review of entomology    February 1, 2017   Volume 62 343-358 doi: 10.1146/annurev-ento-031616-035010
Carpenter S, Mellor PS, Fall AG, Garros C, Venter GJ.African horse sickness virus (AHSV) is a lethal arbovirus of equids that is transmitted between hosts primarily by biting midges of the genus Culicoides (Diptera: Ceratopogonidae). AHSV affects draft, thoroughbred, and companion horses and donkeys in Africa, Asia, and Europe. In this review, we examine the impact of AHSV critically and discuss entomological studies that have been conducted to improve understanding of its epidemiology and control. The transmission of AHSV remains a major research focus and we critically review studies that have implicated both Culicoides and other blood-feeding...
Detection of Schmallenberg virus antibody in equine population of Northern and Northeast of Iran.
Veterinary world    January 18, 2017   Volume 11, Issue 1 30-33 doi: 10.14202/vetworld.2018.30-33
Rasekh M, Sarani A, Hashemi SH.Schmallenberg virus (SBV) is a newly emerging virus in Simbu group that 1 time is reported in 2011 in Germany and now spread to Europe. The clinical signs of infection to this virus are fever, loss of appetite, reduced milk yield and in some cases, diarrhea and in pregnant animals congenital malformations in calves, lambs, and kid goats. Methods: In this study for a serologic survey of SBV, blood samples from 200 horse in different rural areas of the northern and northeast of Iran with the high equine population collected and were analyzed using an indirect ELISA test. Results: Based on our re...
Activity of Culicoides spp. (Diptera: Ceratopogonidae) inside and outside of livestock stables in late winter and spring.
Parasitology research    January 4, 2017   Volume 116, Issue 3 881-889 doi: 10.1007/s00436-016-5361-2
Kameke D, Kampen H, Walther D.Culicoides Latreille, 1809 midge species are the putative vectors of Bluetongue virus (BTV) and Schmallenberg virus (SBV) in Europe. To gain a better understanding of the epidemiology of the diseases, basic knowledge about the overwintering of the vectors is needed. Therefore, we investigated culicoid activity in relation to air temperature at livestock stables during late winter and spring season. Ceratopogonids were captured weekly indoors and outdoors on three cattle farms, three horse farms and one sheep farm in the federal state of Brandenburg, Germany between January and May, 2015 by BG-...
Locally Acquired Eastern Equine Encephalitis Virus Disease, Arkansas, USA.
Emerging infectious diseases    December 15, 2016   Volume 22, Issue 12 2216-2217 doi: 10.3201/eid2212.160844
Garlick J, Lee TJ, Shepherd P, Linam WM, Pastula DM, Weinstein S, Schexnayder SM.No abstract available
Potential vectors of equine arboviruses in the UK.
The Veterinary record    September 30, 2016   Volume 180, Issue 1 19 doi: 10.1136/vr.103825
Chapman GE, Archer D, Torr S, Solomon T, Baylis M.There is growing concern about the increasing risk of disease outbreaks caused by arthropod-borne viruses (arboviruses) in both human beings and animals. There are several mosquito-borne viral diseases that cause varying levels of morbidity and mortality in horses and that can have substantial welfare and economic ramifications. While none has been recorded in the UK, vector species for some of these viruses are present, suggesting that UK equines may be at risk. The authors undertook, therefore, the first study of mosquito species on equine premises in the UK. Mosquito magnet traps and red-bo...
A Serological Protein Microarray for Detection of Multiple Cross-Reactive Flavivirus Infections in Horses for Veterinary and Public Health Surveillance.
Transboundary and emerging diseases    September 15, 2016   Volume 64, Issue 6 1801-1812 doi: 10.1111/tbed.12569
Cleton NB, van Maanen K, Bergervoet SA, Bon N, Beck C, Godeke GJ, Lecollinet S, Bowen R, Lelli D, Nowotny N, Koopmans MPG, Reusken CBEM.The genus Flavivirus in the family Flaviviridae includes some of the most important examples of emerging zoonotic arboviruses that are rapidly spreading across the globe. Japanese encephalitis virus (JEV), West Nile virus (WNV), St. Louis encephalitis virus (SLEV) and Usutu virus (USUV) are mosquito-borne members of the JEV serological group. Although most infections in humans are asymptomatic or present with mild flu-like symptoms, clinical manifestations of JEV, WNV, SLEV, USUV and tick-borne encephalitis virus (TBEV) can include severe neurological disease and death. In horses, infection wi...
Host associations of mosquitoes at eastern equine encephalitis virus foci in Connecticut, USA.
Parasites & vectors    August 30, 2016   Volume 9, Issue 1 474 doi: 10.1186/s13071-016-1765-1
Shepard JJ, Andreadis TG, Thomas MC, Molaei G.Eastern equine encephalitis virus (EEEV) is a highly pathogenic mosquito-borne arbovirus, with active transmission foci in freshwater hardwood swamps in eastern North America, where enzootic transmission is maintained between the ornithophilic mosquito, Culiseta melanura, and wild passerine birds. The role of other locally abundant mosquito species in virus transmission and their associations with vertebrate hosts as sources of blood meals within these foci are largely unknown but are of importance in clarifying the dynamics of enzootic and epidemic/epizootic transmission. Blood-engorged mosqu...
African horse sickness: The potential for an outbreak in disease-free regions and current disease control and elimination techniques.
Equine veterinary journal    July 21, 2016   Volume 48, Issue 5 659-669 doi: 10.1111/evj.12600
Robin M, Page P, Archer D, Baylis M.African horse sickness (AHS) is an arboviral disease of equids transmitted by Culicoides biting midges. The virus is endemic in parts of sub-Saharan Africa and official AHS disease-free status can be obtained from the World Organization for Animal Health on fulfilment of a number of criteria. AHS is associated with case fatality rates of up to 95%, making an outbreak among naïve horses both a welfare and economic disaster. The worldwide distributions of similar vector-borne diseases (particularly bluetongue disease of ruminants) are changing rapidly, probably due to a combination of globalisa...
Evaluation of the Honey-Card Technique for Detection of Transmission of Arboviruses in Florida and Comparison With Sentinel Chicken Seroconversion.
Journal of medical entomology    June 21, 2016   Volume 53, Issue 6 1449-1457 doi: 10.1093/jme/tjw106
Burkett-Cadena ND, Gibson J, Lauth M, Stenn T, Acevedo C, Xue RD, McNelly J, Northey E, Hassan HK, Fulcher A, Bingham AM, van Olphen J, van Olphen A....Zoonotic mosquito-borne viruses, such as the West Nile virus (WNV) and eastern equine encephalitis virus (EEEV), are major public health threats in the United States. Early detection of virus transmission and targeted vector management are critical to protect humans against these pathogens. Sentinel chickens and pool screening of mosquitoes, the most widely used methods of arbovirus early detection, have technical time-lags that compromise their early-detection value. The exploitation of sugar-feeding by trapped mosquitoes for arbovirus surveillance may represent a viable alternative to other ...
Effect of Holding Conditions on the Detection of Chikungunya and Venezuelan Equine Encephalitis Viruses in Mosquito Pools.
Journal of the American Mosquito Control Association    April 23, 2016   Volume 32, Issue 1 51-54 doi: 10.2987/moco-32-01-51-54.1
Andrews ES, Turell MJ.Emerging and re-emerging arboviruses continue to be a threat to global public health, and viral surveillance of mosquito populations is critical for mosquito control operations. Due to the tropical climate of many of the affected areas, it may be difficult to maintain a cold chain as the samples travel from collection sites to laboratories for testing. We determined how suboptimal holding temperatures affected the ability to detect viruses in pools of mosquitoes. Adult female Aedes albopictus and Ae. taeniorhynchus individuals were inoculated with chikungunya virus or Venezuelan equine encepha...
Quantitative Risk Assessment for African Horse Sickness in Live Horses Exported from South Africa.
PloS one    March 17, 2016   Volume 11, Issue 3 e0151757 doi: 10.1371/journal.pone.0151757
Sergeant ES, Grewar JD, Weyer CT, Guthrie AJ.African horse sickness (AHS) is a severe, often fatal, arbovirus infection of horses, transmitted by Culicoides spp. midges. AHS occurs in most of sub-Saharan Africa and is a significant impediment to export of live horses from infected countries, such as South Africa. A stochastic risk model was developed to estimate the probability of exporting an undetected AHS-infected horse through a vector protected pre-export quarantine facility, in accordance with OIE recommendations for trade from an infected country. The model also allows for additional risk management measures, including multiple PC...
Venezuelan Equine Encephalitis Virus Induces Apoptosis through the Unfolded Protein Response Activation of EGR1.
Journal of virology    January 20, 2016   Volume 90, Issue 7 3558-3572 doi: 10.1128/JVI.02827-15
Baer A, Lundberg L, Swales D, Waybright N, Pinkham C, Dinman JD, Jacobs JL, Kehn-Hall K.Venezuelan equine encephalitis virus (VEEV) is a previously weaponized arthropod-borne virus responsible for causing acute and fatal encephalitis in animal and human hosts. The increased circulation and spread in the Americas of VEEV and other encephalitic arboviruses, such as eastern equine encephalitis virus and West Nile virus, underscore the need for research aimed at characterizing the pathogenesis of viral encephalomyelitis for the development of novel medical countermeasures. The host-pathogen dynamics of VEEV Trinidad donkey-infected human astrocytoma U87MG cells were determined by car...
Diversity and ecology survey of mosquitoes potential vectors in Belgian equestrian farms: A threat prevention of mosquito-borne equine arboviruses.
Preventive veterinary medicine    December 29, 2015   Volume 124 58-68 doi: 10.1016/j.prevetmed.2015.12.013
Boukraa S, de La Grandiere MA, Bawin T, Raharimalala FN, Zimmer JY, Haubruge E, Thiry E, Francis F.Emergence of West Nile Virus was recently recorded in several European countries, which can lead to severe health problems in horse populations. Europe is also at risk of introduction of mosquito-borne equine alphavirus from Americas. Prevention of these arboviruses requires a clear understanding of transmission cycles, especially their vectors. To characterize mosquito fauna, their ecology and identify potential vectors of equine arboviruses in Belgium, entomological surveys of six equestrian farms located in the Wolloon Region were conducted during 2011-2012. The harvest of mosquitoes was ba...
Culicoides (Diptera: Ceratopogonidae) and livestock in the Netherlands: comparing host preference and attack rates on a Shetland pony, a dairy cow, and a sheep.
Journal of vector ecology : journal of the Society for Vector Ecology    November 28, 2015   Volume 40, Issue 2 308-317 doi: 10.1111/jvec.12169
Elbers AR, Meiswinkel R.Culicoides (Diptera: Ceratopogonidae) host preferences and attack rates were quantified in early summer at a dairy farm in the Netherlands using livestock tethered at pasture. Midges were aspirated hourly over seven consecutive hours (17:00-23:00) from a dairy cow, a Shetland pony, and a sheep and correspondingly yielded seventeen, thirteen, and nine species. Of the 14,181 midges obtained, approximately 95% belonged to the C. obsoletus complex, C. dewulfi, C. chiopterus, and C. punctatus that together include all proven or potential vectors for arboviral diseases in livestock in northwestern E...
Vector-Host Interactions of Culiseta melanura in a Focus of Eastern Equine Encephalitis Virus Activity in Southeastern Virginia.
PloS one    September 1, 2015   Volume 10, Issue 9 e0136743 doi: 10.1371/journal.pone.0136743
Molaei G, Armstrong PM, Abadam CF, Akaratovic KI, Kiser JP, Andreadis TG.Eastern equine encephalitis virus (EEEV) causes a highly pathogenic mosquito-borne zoonosis that is responsible for sporadic outbreaks of severe illness in humans and equines in the eastern USA. Culiseta (Cs.) melanura is the primary vector of EEEV in most geographic regions but its feeding patterns on specific avian and mammalian hosts are largely unknown in the mid-Atlantic region. The objectives of our study were to: 1) identify avian hosts of Cs. melanura and evaluate their potential role in enzootic amplification of EEEV, 2) assess spatial and temporal patterns of virus activity during a ...