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Topic:Erythrocytes

Erythrocytes, or red blood cells, are the most abundant cell type in the equine circulatory system and are responsible for transporting oxygen from the lungs to tissues throughout the body. They also facilitate the return of carbon dioxide from tissues to the lungs for exhalation. Equine erythrocytes are characterized by their biconcave disc shape, which enhances their flexibility and surface area for gas exchange. The lifespan of erythrocytes in horses is approximately 140-150 days, after which they are removed from circulation and broken down, primarily in the spleen. Variations in erythrocyte count, size, and morphology can indicate various health conditions, including anemia, dehydration, and certain metabolic disorders. This page compiles peer-reviewed research studies and scholarly articles that explore the structure, function, and clinical implications of erythrocytes in equine health.
In vitro activity of ponazuril against Theileria equi.
Veterinary parasitology    November 4, 2011   Volume 185, Issue 2-4 282-285 doi: 10.1016/j.vetpar.2011.10.036
Wise LN, Ueti MW, Kappmeyer LS, Hines MT, White SN, Davis W, Knowles DP.The equid hemoprotozoan parasite Theileria equi is endemic in most regions worldwide. Infection of horses is a cause of significant economic loss due to costs associated with disease and restriction of trade with non-endemic nations. The ability of certain drugs such as imidocarb dipropionate to eliminate persistent T. equi infection and transmission risk is controversial. The anti-protozoal agent ponazuril has been used successfully to treat equine Sarcosystis neurona and Toxoplasma gondii. The hypothesis that ponazuril inhibits replication of T. equi in vitro was tested. T. equi infected equ...
Performance evaluation of the Sysmex pocH-100iV Diff hematology analyzer for analysis of canine, feline, equine, and bovine blood.
Veterinary clinical pathology    November 3, 2011   Volume 40, Issue 4 484-495 doi: 10.1111/j.1939-165X.2011.00372.x
Riond B, Weissenbacher S, Hofmann-Lehmann R, Lutz H.The Sysmex pocH-100iV Diff is an impedance hematology analyzer recently introduced for point-of-care use in veterinary practices in Europe. Objective: The purpose of this study was to validate the pocH-100iV Diff for analysis of blood samples from dogs, cats, horses, and cattle. Methods: Fresh EDTA-blood samples from healthy and ill dogs (115), cats (94), horses (91), and cattle (78) were analyzed on the pocH-100iV Diff and the Cell-Dyn 3500. Results of the automated WBC differential counts were compared with the manual differential counts for 77 dogs, 65 cats, 40 horses, and 46 cattle. HCT we...
Protective effects of passively transferred merozoite-specific antibodies against Theileria equi in horses with severe combined immunodeficiency.
Clinical and vaccine immunology : CVI    October 28, 2011   Volume 19, Issue 1 100-104 doi: 10.1128/CVI.05301-11
Mealey RH, Kappmeyer LS, Ueti MW, Wagner B, Knowles DP.Theileria equi immune plasma was infused into young horses (foals) with severe combined immunodeficiency. Although all foals became infected following intravenous challenge with homologous T. equi merozoite stabilate, delayed time to peak parasitemia occurred. Protective effects were associated with a predominance of passively transferred merozoite-specific IgG3.
Appropriate secondary absolute erythrocytosis in a horse.
The Veterinary record    October 20, 2011   Volume 169, Issue 23 609 doi: 10.1136/vr.100236
Belli CB, Baccarin RY, Ida KK, Fernandes WR.No abstract available
Sequence variations and two levels of MCT1 and CD147 expression in red blood cells and gluteus muscle of horses.
Gene    September 16, 2011   Volume 491, Issue 1 65-70 doi: 10.1016/j.gene.2011.08.030
Koho NM, Mykkänen AK, Reeben M, Raekallio MR, Ilves M, Pösö AR.MCT1-CD147 complex is the prime lactate transporter in mammalian plasma membranes. In equine red blood cells (RBCs), activity of the complex and expression of MCT1 and CD147 is bimodal; high in 70% and low in 30%. We studied whether sequence variations contribute to the bimodal expression of MCT1 and CD147. Samples of blood and cremaster muscle were collected in connection of castration from 24 horses. Additional gluteus muscle samples were collected from 15 Standardbreds of which seven were known to express low amounts of CD147 in RBCs. The cDNA of MCT1 and CD147 together with a promoter regi...
Evaluation of the Mythic 18 hematology analyzer for use with canine, feline, and equine samples. Wassmuth AK, Riond B, Hofmann-Lehmann R, Lutz H.The Mythic 18 is a fully automated hematology bench-top analyzer using impedance technology for a complete blood cell count (CBC) and a 3-part white blood cell count (WBC) differential. The purpose of the current study was to evaluate the Mythic for assessment of agreement, precision, linearity, carry-over, stability, and usability under practice conditions. Ethylenediamine tetra-acetic acid-blood samples from 122 dogs, 140 cats, and 123 horses were analyzed with the Mythic and reference methods (Sysmex XT-2000iV, manual hematocrit, and microscopic WBC differentiation). Pearson's coefficient o...
Transplacental transmission of Babesia equi (Theileria equi) from carrier mares to foals.
Journal of parasitic diseases : official organ of the Indian Society for Parasitology    September 3, 2011   Volume 36, Issue 1 31-33 doi: 10.1007/s12639-011-0072-1
Chhabra S, Ranjan R, Uppal SK, Singla LD.This article communicates the first per-acute and fatal clinical report of transplacental transmission of Theileria equi from carrier mothers to foals from Punjab, India. Two cases of equine piroplasmosis due to T. equi were diagnosed in neonatal foals borne to Throughbred mares. High degree of parasitemia with annular and maltese cross shaped parasites in erythrocytes was seen in both cases. Blood cellular changes revealed leucopenia and neutropenia with mild degenerative left shift. Anaemia was macrocytic normochromic type. Intense yellow discoloration of mucous membranes indicated jaundice....
Identification of a mutation associated with fatal Foal Immunodeficiency Syndrome in the Fell and Dales pony.
PLoS genetics    July 7, 2011   Volume 7, Issue 7 e1002133 doi: 10.1371/journal.pgen.1002133
Fox-Clipsham LY, Carter SD, Goodhead I, Hall N, Knottenbelt DC, May PD, Ollier WE, Swinburne JE.The Fell and Dales are rare native UK pony breeds at risk due to falling numbers, in-breeding, and inherited disease. Specifically, the lethal Mendelian recessive disease Foal Immunodeficiency Syndrome (FIS), which manifests as B-lymphocyte immunodeficiency and progressive anemia, is a substantial threat. A significant percentage (∼10%) of the Fell ponies born each year dies from FIS, compromising the long-term survival of this breed. Moreover, the likely spread of FIS into other breeds is of major concern. Indeed, FIS was identified in the Dales pony, a related breed, during the course of t...
Effects of chronic acetazolamide administration on gas exchange and acid-base control in pulmonary circulation in exercising horses.
Equine veterinary journal. Supplement    May 28, 2011   Issue 38 40-50 doi: 10.1111/j.2042-3306.2010.00240.x
Vengust M, Stämpfli H, De Moraes AN, Teixeiro-Neto F, Viel L, Heigenhauser G.Carbonic anhydrase (CA) catalyses the hydration/dehydration reaction of CO(2) and increases the rate of Cl(-) and HCO(3)(-) exchange between the erythrocytes and plasma. Therefore, chronic inhibition of CA has a potential to attenuate CO(2) output and induce greater metabolic and respiratory acidosis in exercising horses. Objective: To determine the effects of Carbonic anhydrase inhibition on CO(2) output and ionic exchange between erythrocytes and plasma and their influence on acid-base balance in the pulmonary circulation (across the lung) in exercising horses with and without CA inhibition....
Oral supplementation with superoxide dismutase in Standardbred trotters in training: a double-blind placebo-controlled study.
Equine veterinary journal. Supplement    May 27, 2011   Issue 38 375-381 doi: 10.1111/j.2042-3306.2010.00266.x
Notin C, Vallon L, Desbordes F, Leleu C.Intense physical exercise produces an excess of reactive oxygen species which can disturb the antioxidant/oxidant balance of the horse in training. Several classes of antioxidant dietary compounds have been suggested to provide health benefits and there is evidence that consumption of these products leads to a reduction in the expression of various pro-inflammatory and/or oxidative stress biomarkers. The recent development of a new galenic system allows the oral delivery of the antioxidant enzyme: superoxide dismutase (SOD). This has been developed from a specific melon variety with a particul...
Effects of intravenous aminocaproic acid on exercise-induced pulmonary haemorrhage (EIPH).
Equine veterinary journal. Supplement    May 27, 2011   Issue 38 256-260 doi: 10.1111/j.2042-3306.2010.00247.x
Buchholz BM, Murdock A, Bayly WM, Sides RH.The antifibrinolytic, 6-aminohexanoic acid, also named aminocaproic acid (ACA), has been used empirically as a treatment for exercise-induced pulmonary haemorrhage (EIPH) on the unsubstantiated basis that transient coagulation dysfunction may contribute to its development. Objective: To assess the effect of ACA on bronchoalveolar lavage fluid (BALF) erythrocyte counts in horses performing treadmill exercise at an intensity greater than that needed to reach maximal oxygen consumption. Methods: Eight Thoroughbreds were exercised to fatigue 3 times on a 10% inclined treadmill at a speed for which...
Experimental mild pulmonary inflammation promotes the development of exercise-induced pulmonary haemorrhage.
Equine veterinary journal. Supplement    May 27, 2011   Issue 38 235-239 doi: 10.1111/j.2042-3306.2010.00295.x
McKane SA, Slocombe RF.Histological studies of exercise-induced pulmonary haemorrhage (EIPH) reveals inflammatory infiltrates within areas of lung that contain haemorrhage. This has resulted in the hypothesis that pulmonary inflammation could cause EIPH or contribute to an increased risk or severity of EIPH. Objective: To determine whether experimentally-induced pulmonary inflammation predisposes the lung to haemorrhage during exercise, by evaluating the bronchoalveolar lavage (BAL) cytology of normal and inflamed regions of lung following high speed treadmill exercise. Methods: Transendoscopic airway inoculations o...
Changes in arterial, mixed venous and intraerythrocytic ion concentrations during prolonged exercise.
Equine veterinary journal. Supplement    May 27, 2011   Issue 38 185-190 doi: 10.1111/j.2042-3306.2010.00189.x
Meyer ND, Bayly WM, Sides RH, Wardrop KJ, Slinker BK.Prolonged equine exercise can cause hypochloraemic alkalosis and hypokalaemia secondary to the loss of hypertonic sweat. Movement of ions in and out of erythrocytes during exercise may help regulate acid-base balance and changes in plasma ion concentrations. The extent to which this happens during prolonged equine exercise has not been reported. Objective: To measure changes in blood gases and major plasma and intraerythrocytic (iRBC) ion concentrations of horses undergoing prolonged submaximal exercise. Methods: Six horses were trotted at ∼ 30% VO2max on a treadmill for 105 min. Arterial ((...
Expression of lactate transporters MCT1, MCT2 and CD147 in the red blood cells of three horse breeds: Finnhorse, Standardbred and Thoroughbred.
Equine veterinary journal. Supplement    May 27, 2011   Issue 38 161-166 doi: 10.1111/j.2042-3306.2010.00205.x
Mykkänen AK, Pösö AR, McGowan CM, McKane SA.In exercising horses, up to 50% of blood lactate is taken up into red blood cells (RBCs). Lactate transporter proteins MCT1, MCT2 and CD147 (an ancillary protein for MCT1) are expressed in the equine RBC membrane. In Standardbreds (SB), lactate transport activity is bimodally distributed and correlates with the amount of MCT1 and CD147. About 75% of SB studied have high lactate transport activity in RBCs. In other breeds, the distribution of lactate transport activity is unknown. Objective: To study whether similar bimodal distribution of MCT1 and CD147 is present also in the racing Finnhorse ...
The role of RTX toxins in host specificity of animal pathogenic Pasteurellaceae.
Veterinary microbiology    May 19, 2011   Volume 153, Issue 1-2 51-58 doi: 10.1016/j.vetmic.2011.05.018
Frey J.RTX toxins are bacterial pore-forming toxins that are particularly abundant among pathogenic species of Pasteurellaceae, in which they play a major role in virulence. RTX toxins of several primary pathogens of the family of Pasteurellaceae are directly involved in causing necrotic lesions in the target organs. Many RTX toxins are known as haemolysins because they lyse erythrocytes in vitro, an effect that is non-specific, but which serves as a useful marker in bacteriological identification and as an easily measurable signal in vitro in experimental studies. More recent studies have shown that...
In-vitro hypocoagulability on whole blood thromboelastometry associated with in-vivo expansion of red cell mass in an equine model.
Blood coagulation & fibrinolysis : an international journal in haemostasis and thrombosis    April 22, 2011   Volume 22, Issue 5 424-430 doi: 10.1097/MBC.0b013e3283464f83
McMichael M, Smith SA, McConachie EL, Lascola K, Wilkins PA.In several species, there is a strong correlation between indicators of red cell mass (RCM) and thromboelastometry results. The horse has a reliable, temporary, polycythemia in response to phenylephrine infusion. Our objective was to evaluate the effects of an in-vivo increase in circulating RCM on thromboelastometry results in an equine model of transient polycythemia. Six healthy research horses had whole blood thromboelastometry with contact activator and tissue factor initiation after recalcification of citrated samples. Additional samples were frozen for thrombin-antithrombin (TAT). Compl...
Failure of the Amblyomma cajennense nymph to become infected by Theileria equi after feeding on acute or chronically infected horses.
Experimental parasitology    April 9, 2011   Volume 128, Issue 4 324-327 doi: 10.1016/j.exppara.2011.03.016
Ribeiro MF, da Silveira JA, Bastos CV.Tick-borne diseases in horses are caused by the intraerythrocytic protozoan parasites Theileria equi and Babesia caballi. Although T. equi is highly endemic in Latin America, the New World vector of this important parasite is controversial. The aim of this study was to test the ability of nymph Amblyomma cajennense ticks acquire infection by T. equi following feeding on infected horses. Three experiments were performed: tick acquisition of T. equi from an experimentally infected horse, tick acquisition of T. equi from naturally infected foals and tick acquisition of T. equi from a chronically ...
Disturbance of oxidant/antioxidant equilibrium in horses naturally infected with Trypanosoma evansi.
Veterinary parasitology    March 31, 2011   Volume 180, Issue 3-4 349-353 doi: 10.1016/j.vetpar.2011.03.029
Ranjithkumar M, Kamili NM, Saxena A, Dan A, Dey S, Raut SS.Oxidant/antioxidant equilibrium disturbance has already been reported in trypanosome infections by several authors. The present study was aimed to explore the possible oxidant/antioxidant disturbance in surra of naturally infected horses before and after treatment. Fifteen naturally infected horses were chosen to analyse erythrocytic indices, platelet counts, lipid peroxides (LPO), nitric oxide (NO), reduced glutathione (GSH), superoxide dismutase (SOD) and catalase (CAT) while six healthy animals acted as control. There was a highly significant (P<0.001) reduction in red blood cell (RBC) c...
Production of recombinant EMA-1 protein and its application for the diagnosis of Theileria equi using an enzyme immunoassay in horses from São Paulo State, Brazil. Baldani CD, Hilario E, Nakaghi AC, Bertolini MC, Machado RZ.The erythrocytic-stage surface protein, Equi Merozoite Antigen 1 (EMA-1), is a major candidate for the development of a diagnostic antigen for equine piroplasmosis. In order to establish an effective diagnostic method for practical use, the gene encoding the entire EMA-1 of Theileria equi Jaboticabal strain was cloned and expressed in Escherichia coli as a histidine-tagged protein (His6-EMA1). The expressed EMA-1 reacted with specific antibodies in Western blot and had an apparent molecular mass of 34 kDa which was largely consistent with its theoretical value. The nucleotide sequence of the E...
Prevalence of Theileria equi and Babesia caballi infection in horses from northern Italy.
Vector borne and zoonotic diseases (Larchmont, N.Y.)    January 23, 2011   Volume 11, Issue 7 955-956 doi: 10.1089/vbz.2010.0193
Grandi G, Molinari G, Tittarelli M, Sassera D, Kramer LH.Babesia caballi and Theileria equi are the causative agents of equine piroplasmosis. In this epidemiological study, 294 horses reared in a rural area of northern Italy were studied. During January 2008-January 2009, blood samples were taken for serology (indirect fluorescent antibody test) and for polymerase chain reaction (PCR). Data on the geographical area, sex, and age were collected for statistical analysis of risk factors associated with infection. A seroprevalence of 8.5% was found: 8.2% of the animals were positive for anti-T. equi antibodies and 0.3% for anti-B. caballi antibodies. No...
Na+, K(+)-ATPase activity and ATP concentration in horses of the Wielkopolski breed in relation to age.
Polish journal of veterinary sciences    January 1, 2011   Volume 14, Issue 4 635-642 doi: 10.2478/v10181-011-0094-8
Suska M, Brucka-Jastrzebska E, Kawczuga D.This study aimed at determining relationships between the age of the Wielkopolski horses, ATP in whole blood and in the erythrocytes, and between erythrocyte Na+, K(+)-ATPase activity, and serum concentrations of mineral components. ATP was measured in whole blood and in erythrocytes by HPLC method. Serum concentrations of Ca2+ and Mg2+ were measured spectrophotometrically, while Na+ and K+ by flame photometry. In horses aged from 4 to 48 months, a dynamic decrease in ATP activity was found. Erythrocyte Na+, K(+)-ATPase activity decreased proportionally with the decrease in ATP activity. The r...
Occurrence of Theileria equi in horses raised in the Jaboticabal microregion, São Paulo State, Brazil. Baldani CD, Nakaghi AC, Machado RZ.Blood and serum samples from 170 horses raised in the Jaboticabal microregion, São Paulo State, Brazil, were collected and tested by microscopic examination of blood smears, indirect fluorescent antibody test (IFAT) and nested polymerase chain reaction (nPCR) for Theileria equi infections. The association among the test results was verified by the McNemar test. During the examination of thin blood smears, parasites were detected in six (3.52%) horses. Anti-T. equi antibodies were detected in 100% sera samples, with titers ranging between 1:80 and 1:5120. The nPCR based on the T. equi merozoit...
Effects induced by exercise on lymphocyte β-adrenergic receptors and plasma catecholamine levels in performance horses.
Research in veterinary science    December 17, 2010   Volume 92, Issue 1 116-120 doi: 10.1016/j.rvsc.2010.11.002
Cuniberti B, Badino P, Odore R, Girardi C, Re G.The effect of dynamic exercise on complete blood cell count, lymphocyte β-adrenergic receptor and plasma catecholamine (adrenaline and noradrenaline) levels in horses performing different disciplines were investigated during rest and after exercise. Blood samples were collected from jumping horses (n=6), Arabian Endurance horses (n=6) and Standardbred trotters (n=6) before and immediately after competition. Dynamic exercise caused a significant increase in red blood cell count (Standardbred trotters: P=0.0012), haemoglobin concentration (jumping horses: P=0.001; Standardbred trotters: P=0.01)...
A case of transplacental transmission of Theileria equi in a foal in Trinidad.
Veterinary parasitology    October 20, 2010   Volume 175, Issue 3-4 363-366 doi: 10.1016/j.vetpar.2010.10.019
Georges KC, Ezeokoli CD, Sparagano O, Pargass I, Campbell M, D'Abadie R, Yabsley MJ.Equine piroplasmosis due to Theileria equi and Babesia caballi is endemic in Trinidad. A case of equine piroplasmosis due to T. equi was diagnosed in a thoroughbred foal at 10h post-partum. A high parasitaemia (63%) of piroplasms was observed in a Wright-Giemsa(®) stained thin blood smear from the foal. In addition, the 18S rRNA gene for Babesia/Theileria was amplified from DNA extracted from the blood of the foal and the mare. Amplified products were subjected to a reverse line blot hybridization assay (RLB), which confirmed the presence of T. equi DNA in the foal. The mare was negative by R...
Morphological and biochemical changes in the blood of horses naturally infected with Gasterophilus sp. larvae.
Polish journal of veterinary sciences    August 25, 2010   Volume 13, Issue 2 247-251 
Pawlas-Opiela M, Sołtysiak Z, Gorczykowski M.Gasterophilus sp. constitute a group of specific parasites, which larval forms are found in horses and other phylogenetically related species--donkeys, mules and zebras. Their invasion is associated with marked fluctuations of hematological and biochemical blood parameters, including the activity of many enzymes and the electrolyte concentration. The purpose of the study was to analyze changes in selected biochemical and morphological blood parameters of horses infected with botfly larvae. The experimental group was formed of horses infested by Gasterophilus sp. larvae whereas the control grou...
Pregnancy influences the hematological profile of Carthusian broodmares.
Polish journal of veterinary sciences    August 25, 2010   Volume 13, Issue 2 393-394 
Satué K, Muñoz A, Blanco O.In order to investigate the effect of pregnancy on equine hematological profile, venous blood samples were taken from 44 Carthusian broodmares every two weeks during pregnancy, divided into three periods: I (conception to 110 days), II (111-220 days) and III (222 days to delivery). Routine hematological analysis was performed. Packed cell volume (PCV) increased in period II probably as a consequence of the climatic conditions or as a result of physical stress of lactation during period I. After that, a reduction in PCV was found but this was not matched with a similar decrease in total serum p...
Blood cells RNA biomarkers as a first long-term detection strategy for EPO abuse in horseracing.
Drug testing and analysis    August 6, 2010   Volume 2, Issue 7 339-345 doi: 10.1002/dta.146
Bailly-Chouriberry L, Noguier F, Manchon L, Piquemal D, Garcia P, Popot MA, Bonnaire Y.Recombinant human erythropoietins (rHuEPOs) are glycoproteins drugs, produced by the pharmaceutical industry to restore production of red blood cells by stimulating human bone marrow for which this pathology has been diagnosed. It is suspected that these molecules are diverted as doping agents in horseracing to enhance oxygen transport and aerobic power in racehorses. Although indirect double-blotting or direct liquid chromatography-mass spectrometry (LC-MS) methods have been developed to confirm the presence of rHuEPO in a sample, the short detection time (48 h) is still a problem for doping ...
Use of an in vitro biotinylation technique for determination of posttransfusion survival of fresh and stored autologous red blood cells in Thoroughbreds.
American journal of veterinary research    August 3, 2010   Volume 71, Issue 8 960-966 doi: 10.2460/ajvr.71.8.960
Owens SD, Johns JL, Walker NJ, Librach FA, Carrade DD, Tablin F, Borjesson DL.To evaluate N-hydroxysuccinimide (NHS)-biotin labeling of equine RBCs and determine posttransfusion survival of autologous equine RBCs stored in citrate phosphate dextrose adenine-1 (CPDA-1) for 0, 1, 14, and 28 days. Methods: 13 healthy adult Thoroughbreds. Methods: Serial dilutions of biotin and streptavidin-phycoerythrin (PE) were evaluated in vitro in blood collected from 3 horses. One horse was used to determine RBC distribution and recovery. Twelve horses were allocated to 4 groups for in vivo experiments in which blood was collected into CPDA-1. Blood was labeled with biotin and reinfus...
Low-molecular-weight heparin dosage in newborn foals.
Journal of veterinary internal medicine    July 28, 2010   Volume 24, Issue 5 1190-1195 doi: 10.1111/j.1939-1676.2010.0568.x
Armengou L, Monreal L, Delgado MÁ, Ríos J, Cesarini C, Jose-Cunilleras E.Heparin is used in humans as prophylaxis of hypercoagulable states and disseminated intravascular coagulation (DIC). However, babies need a higher heparin dose than do adults. Septic neonate foals are at high risk of hypercoagulable state and DIC, and there is limited objective information about heparin dose for equine neonates. Objective: To assess whether neonate foals require higher dosages of low-molecular-weight heparin (LMWH) than adults. Methods: Eighteen healthy and 11 septic neonate foals. Methods: Experimental and clinical studies. Firstly, healthy foals were randomly distributed in ...
Sequence heterogeneity in the equi merozoite antigen gene (ema-1) of Theileria equi and development of an ema-1-specific TaqMan MGB assay for the detection of T. equi.
Veterinary parasitology    April 28, 2010   Volume 172, Issue 1-2 33-45 doi: 10.1016/j.vetpar.2010.04.025
Bhoora R, Quan M, Matjila PT, Zweygarth E, Guthrie AJ, Collins NE.Although a quantitative real-time PCR assay (qPCR) assay for the detection of Theileria equi has been developed and evaluated, it is possible that additional, as yet undetected 18S rRNA gene sequence variants may exist. A qPCR assay targeting a different gene, used in conjunction with the T. equi 18S rRNA qPCR assay, could assist in the detection of all T. equi genotypes in field samples. A T. equi ema-1-specific qPCR (Ueti et al., 2003) was tested on 107 South African field samples, 90 of which tested positive for T. equi antibody using the immuno-fluorescent antibody test (IFAT). The qPCR as...
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