Trotting Horses refers to a specific gait in equines characterized by a two-beat diagonal stride, where the legs move in pairs diagonally across the horse's body. This gait is commonly observed in various horse breeds and is a fundamental aspect of disciplines such as harness racing and dressage. Trotting is often evaluated for its rhythm, balance, and speed, particularly in competitive settings. This topic encompasses a range of research studies and scholarly articles that investigate the biomechanics, training techniques, and performance metrics associated with trotting in horses. Additionally, the page includes research on the physiological and anatomical adaptations that enable this specific gait, as well as its implications for equine health and athletic performance.
Clayton HM.High speed cinematography was used to measure the timing and distance characteristics of the trot in five sound horses. Two farriery treatments were compared: (1) normal, in which the dorsal hoof wall was aligned with the pastern axis in all four feet and (2) acute hind hoof angulation, in which the hind toes were allowed to grow relatively longer than the heels, so that the angulation was approximately 10 degrees lower than the normal hind hoof angle for the individual animal. Comparison between the two treatments showed that the acute hind hoof angulation was associated with significant incr...
van Weeren PR, van den Bogert AJ, Barneveld A.Skin displacement was investigated at 16 sites in the front and hind limbs of four Dutch Warmblood horses at the trot. For visualisation of the bone under the skin, a measuring device mounted on a Steinmann pin driven into the bone under investigation, was employed. Mean displacements ranged from 8 mm (X-displacements proximal and distal metacarpus) to 142 mm (Y-displacement caudal part of greater trochanter). When compared with previously determined skin displacement patterns at walk, mean displacements at trot were of the same magnitude whereas the shape of the time-displacement curve was di...
Clayton HM.High speed cinematography was used to describe the stride kinematics in a group of six, sound horses subjected to two farriery treatments, designated 'normal' and 'acute' hoof angulation. Normal was defined as having the dorsal hoof wall aligned with the pastern axis, whereas the acute hoof angulation was approximately 10 degrees lower than the normal angle for the individual horse. The acute angle was achieved by allowing the toes to grow relatively longer than the heels. The results of a multivariate analysis of variance showed significant differences between the two farriery treatments in t...
Art T, Desmecht D, Amory H, Lekeux P.The aim of the present experiment was to investigate the hypothesis that there is a respiration-locomotion coupling in trotting equines. Therefore the respiratory airflow (V), the gastric pressure (Pga) and pleural pressure (Ppl) changes and the step frequency (SF) were simultaneously recorded in 5 trotting ponies (body weight: 255 +/- 15 kg; age: 3.5 +/- 0.4 years). Airflow was measured using a Fleisch pneumotachograph fixed on a face mask. Esophageal and gastric balloon catheters coupled to pressure transducers allowed the recording of Ppl and Pga. The exercise was performed on a treadmill a...
Pösö AR, Viljanen-Tarifa E, Soveri T, Oksanen HE.Effects of graded intensity exercise on plasma lipids was studied in Standardbred and Finnishbred Trotters. The rate of lipolysis indicated by the elevated plasma concentrations of glycerol increased parallel with the intensity of the trot in the Standardbreds, but not as clearly in the Finnishbred trotters. During the exercise plasma triglyceride concentration increased significantly and the increase correlated with the intensity of the exercise as well as the activity of lipolysis. Together with the increase in plasma triglycerides, there was a parallel increase in the pre-beta fraction of l...
Gottlieb M, Essén-Gustavsson B, Lindholm A, Persson SG.Five Standardbred trotters were trained on a treadmill 3 times/wk for 12 wk by intervals of draft-loaded exercise. The draft load was 34 kp and the velocity approximately 7 m/s. Muscle biopsies were taken from the gluteus medius and longissimus muscles before training and after 2, 4, 8, and 12 wk of training and from the brachiocephalicus muscle before and after training. Both the percentage and the area of type IIa fibers increased and the percentage of type IIb fibers decreased in the gluteus medius muscle during the first 2 wk of training, and then no further significant difference was note...
Andersson-Eklund L, Andersson L, Sandberg K.A study was carried out to attempt to explain the basis of the association between the Es locus and starting proportion in Swedish Trotters which had been observed previously. The effect of Es genotype on starting proportion has diminished among horses born in the late 1970s. There are indications that the incidence of leg lesions varies between Es genotypes. If this is true, it is possible that the decreasing effect of the Es locus on starting proportion is due to the environmental changes which have been made at race tracks in the early 1980s to reduce the strain on the legs of the trotters....
Kobluk CN, Schnurr D, Horney FD, Sumner-Smith G, Willoughby RA, Dekleer V, Hearn TC.High-speed cinematography with computer aided analysis was used to study equine hindlimb kinematics. Eight horses were filmed at the trot or the pace. Filming was done from the side (lateral) and the back (caudal). Parameters measured from the lateral filming included the heights of the tuber coxae and tailhead, protraction and retraction of the hoof and angular changes of the tarsus and stifle. Abduction and adduction of the limb and tarsal height changes were measured from the caudal filming. The maximum and minimum values plus the standard deviations and coefficients of variations are prese...
Kiper ML, MacAllister C, Qualls C.A cranial mesenteric arterial aneurysm with fistulous tracts to the cecum and ileum was discovered in a Missouri Fox Trotter stallion. The principal clinical sign was hematochezia, observed at 7- and 10-day intervals. The lesion may have been attributable to arteritis caused by fourth-stage larvae of Strongylus vulgaris.
Gottlieb M, Essén-Gustavsson B, Lindholm A, Persson SG.Circulatory and muscle metabolic responses were studied in 10 horses which all performed incremental draught work at a low trotting speed on a treadmill (D-test) and also exercise with gradually increasing velocities (S-test). Exercise was continued until the horses could no longer maintain the weights above the floor or maintain speed trotting without changing gait to a gallop. Muscle biopsies were taken from the gluteus and the semitendinosus muscles before, and immediately after, exercise. The heart rate (HR) increased linearly with both increasing draught resistance and velocity and reache...
Weitkamp LR, MacCluer JW, Guttormsen SA, King RH.The transmission ratios of alleles at 12 protein marker loci were computed individually for American Standardbred stallions in a genealogy of 5392 phenotyped horses. Over all loci there was significant gene transmission distortion for trotting stallions (p = 0.0019) but not for pacing stallions (p = 0.99). The transmission distortion was due to sire-specific effects (p = 0.0024) and not to increased transmission of one or the other allele of a given heterozygous genotype (p = 0.21). Individual-specific, non-random transmission of homologous chromosomes may provide a mechanism for selection to ...
MacCluer JW, Bailey E, Weitkamp LR, Blangero J.We have analysed the effects of ELA alleles and sire-dam ELA incompatibility on two measures of fertility, gestation length and foaling rate, in American Standardbred horses. Using multivariate statistical methods, we corrected for the effects of confounding factors such as dam and sire age, parity, inbreeding, and sire-dam kinship. These analyses revealed substantial differences between Standardbred trotters and pacers in the effects of several confounding factors. There appear to be no ELA effects on gestation length in either trotters or pacers. However our results suggest that there may be...
Pösö AR, Soveri T, Alaviuhkola M, Lindqvist L, Alakuijala L, Mäenpää PH, Oksanen HE.Previous studies in humans have shown that alanine is released from the skeletal muscle in proportion to the work load. We have measured plasma alanine and urea concentrations in well-trained Standardbred and Finnish-bred (cold-blooded) trotters after a graded-intensity exercise and during recovery to study metabolic responses to exercise in this animal model. As controls we measured blood lactate, pyruvate, and glucose concentrations as well as hematocrit values. Metabolic responses to exercise were closely reflected in all these parameters. Plasma alanine increased relatively more than plasm...
Drevemo S, Fredricson I, Hjertén G, McMiken D.Ten trotting Standardbred colts were recorded by high-speed cinematography at the ages of eight, 12 and 18 months. The horses were trotting on a treadmill operating at 4.0 m/secs. Five horses were subjected to a programme of intensified training from eight months of age, whereas the others were not trained and acted as controls. The films were analysed on a semi-automatic film-reading equipment and a number of variables used to demonstrate the gait symmetry were calculated and scaled by computer. Certain differences between left and right diagonal and contralateral pair of limbs, respectively,...
Cothran EG, MacCluer JW, Weitkamp LR, Bailey E.American Standardbred horses are divided into two groups based upon gait: the trot and the pace. The tendency to trot (diagonally opposite legs moving forward together) or pace (the two legs on the same side of the body moving forward together) appears to be genetically determined, although no formal genetic analysis has been undertaken. There is nearly complete assortative mating for gait; however, about 20% of the offspring sired by trotters are registered as pacers, while fewer than 1% of those sired by pacers are registered as trotters. Electrophoretically detectable genic variation at 13 ...
Valberg S.The glycogen depletion patterns in the gluteus muscle of Standardbred horses were studied under different trotting intensities. After racing significant glycogen depletion was found in all Type I and IIA fibres and in a varying percentage of Type IIB fibres, depending on the individual horse. When horses performed exercise over a short distance (5 to 8 km) at three different speeds, glycogen depletion was difficult to detect for the faster speeds (10 and 8 m/sec) but notably involved a major proportion of Type I fibres at the slowest speed (6 m/sec). When exercise was prolonged over a greater ...
Hodges AA, Harrison AG, Wathes CM.The heights of 89 horses were measured at the withers before and after half a furlong of trotting exercise. The mean (+/- sd) height increase after exercise was 1.75 +/- 0.86 cm and the horses returned to their resting height within seven minutes. There was no linear relationship between gain in height and pre-exercise height.
Wintzer HJ.A long-term treatment with biotin (vitamin H) in 5 warm-blooded horses and 10 trotter horses is reported. The dose of 0.031-0.037 mg/kg body weight was well tolerated, and with a therapy period up to 10 months an improvement of the horn quality of the growing hoof could be attained as it had not been possible before with other measures. Biotin (Gabiotan) is recommendable as a therapeutic in all cases of hoof problems which are based on disturbed horn elasticity.
Orton RK, Hume ID, Leng RA.Rates of passage of fluid and particulate digesta markers and apparent digestibility estimated by three methods were compared in yearling horses fed high (14 per cent) or low (8 per cent) crude protein diets with one of two levels of exercise (0 or 12 km trotting per day at 12 km/h). Mean retention times (MRT) of the fluid marker (51Cr-EDTA) were shorter than those of the particulate marker (ruthenium-phenanthroline). There were no significant effects of dietary protein level on passage of either marker or on apparent digestibility of dry matter. Exercise increased voluntary feed intake and ap...
Ratzlaff MH, Shindell RM, DeBowes RM.Blood pressures from the catheterized lateral digital vein of the fore-limbs of 6 clinically normal horses were measured at rest, at the walk, and at the trot. Digital venous pressures were compared with the phases of the stride and weight-bearing forces, using electrogoniometry and a force platform. Rapid increases in digital venous pressures to maximal values were observed immediately before maximal forces during the support period of the stride. At the trot, increases in peak vertical forces were paralleled by increases in peak digital venous pressures. Seemingly, the hydrodynamics of the d...
Ingvast-Larsson C, Paalzow G, Paalzow L, Ottosson T, Lindholm A, Appelgren LE.The pharmacokinetics of theophylline were determined in Standardbred trotters after single intravenous and oral administration. A bi-exponential equation was fitted to the intravenous data and a tri-exponential equation to the oral data. The biological half-life of theophylline was found to be 14.8 h, the volume of distribution 1.02 l/kg and the total plasma clearance 0.86 ml/kg/min. The oral absorption of the drug was complete (bioavailability 108%) and rapid (absorption half-life 0.4 h).
Sloet van Oldruitenborgh-Oosterbaan MM, van den Ingh TS.A trotter stallion showing symptoms of emaciation was suspected of disease of the liver associated with cholestasis in view of clinical symptoms (poor appetite, sluggishness, jaundice and oedema) and the results of examination of the blood (increased concentrations of gamma-glutamyl, transpeptidase, sorbitol dehydrogenase, alkaline phosphatase, aspartate aminotransferase and markedly increased conjugated bilirubin). A specimen removed at biopsy of the liver revealed the presence of portal fibrosis and severe cholestasis. At autopsy, it was found that very extensive fibrosis of the pancreas (pr...
Varewyck H, Bouquet Y, Lazary S, Guérin G, Van de Weghe A, Van Zeveren A.158 Belgian Saddlebreds, 130 Belgian Trotters, 108 Belgian Draft horses and 92 Shetland ponies have been typed for serologically defined antigens at the ELA and ELY systems. Gene frequencies were estimated in each breed for the internationally established ELA, ELY-1 and ELY-2 alleles as well as for locally assigned additional ELA markers and for subtypes of ELA-W3, W9 and W11. The distribution of ELA alleles was in agreement with the expected Hardy-Weinberg equilibrium for the 4 horse breeds described here. Differences in gene frequencies between these main Belgian horse populations were obser...
Alitalo I, Kärkkäinen M.Ataxia caused by a focal compression of the cervical spinal cord was diagnosed in four young standardbred trotting horses. Diagnosis was verified by myelography. Changes in the cervical vertebral column were studied using microradiographic and histologic methods. In the vertebrae involved, there was irregularity of the cartilaginous growth zone, cracks with a loose fragment and disturbance in the enchondral ossification. These changes resemble osteochondrosis. Porous appearance in lateral and ventral funiculi as well as mural calcified plaques in the small vessels of the white matter were foun...
van den Hoven R.Venous lactate concentrations were determined in standard-bred trotters following exercise of varying intensity to obtain a useful parameter in determining the working capacity of horses. After standard exercise, well-trained horses produced less lactate than did inadequately trained horses. During the post-exercise resting period, lactate disappeared faster from the blood in well-trained horses and young lightly trained horses than it did in poorly trained horses. Steady-state training resulted in lower lactate levels than those recorded after racing, whereas interval training resulted in the...
Hartikka P, Dahlbom M, Westermarck H.In four training stables five trotter horses of the Finnish coldblooded breed and seven warmblooded trotters were treated with a ferrissaccharin preparation (Hippiron, Hausmann) intravenously. A short increase in heart frequency after the injection was noted. The cold solution injections, less than + 15 degrees C, may cause dangerous reactions. All horses showed an enhanced appetite, a substantially livier habitus after treatment.
Wintzer HJ.A long-term treatment with biotin (vitamin H) in 5 warm-blooded horses and 10 trotter horses is reported. The dose of 0.031-0.037 mg/kg body weight was well tolerated, and with a therapy period up to 10 months an improvement of the horn quality of the growing hoof could be attained as it had not been possible before with other measures. Biotin (Gabiotan) is recommendable as a therapeutic in all cases of hoof problems which are based on disturbed horn elasticity.
Grauerholz H, Jaeschke G.After the onset of training ("breaking") in young standardbred horses within continuing investigations changes of vectors in the QRS-complex were considered. These changes accomplished in two stages. Mainly within the first three months of training the vector R (VR) in limb leads changed to the left and to caudal and dorsal. In a second interval between five and seven months after breaking the vector S (VS) changed to cranial. As main cause the different character of training in both phases is discussed. Additional indicators of age- or development-related causes were not found.
Ambros'eva ED, Khokhriakova ZhA, Glazko VI.The genetic structures of Orlov's and Russian trotters for 16 biochemical systems have been comparatively analyzed. Polymorphism of 6 systems of these horse groups is revealed. The main genetic differentiation between Orlov's and Russian trotters is observed on the transferrin and phosphoglucomutase loci.
Hodges AA, Harrison AG, Wathes CM.The heights of 89 horses were measured at the withers before and after half a furlong of trotting exercise. The mean (+/- sd) height increase after exercise was 1.75 +/- 0.86 cm and the horses returned to their resting height within seven minutes. There was no linear relationship between gain in height and pre-exercise height.
Preedy DF, Colborne GR.Metabolic and mechanical energy costs of locomotion can be combined to calculate locomotor efficiency, which is the quotient of the mechanical energy and metabolic costs. The purpose of this pilot study was to evaluate the mechanical and metabolic energy costs of locomotion at a range of 7 trotting speeds (2.5 to 6.2 m/s) on a level treadmill. A single, sound Thoroughbred horse was modelled as a system of 15 linked segments incorporating all 4 limbs, head, neck and trunk. The horse performed a continuous incremental exercise test at increasing trotting speeds while VO2 was recorded using a bre...
Hartikka P, Dahlbom M, Westermarck H.In four training stables five trotter horses of the Finnish coldblooded breed and seven warmblooded trotters were treated with a ferrissaccharin preparation (Hippiron, Hausmann) intravenously. A short increase in heart frequency after the injection was noted. The cold solution injections, less than + 15 degrees C, may cause dangerous reactions. All horses showed an enhanced appetite, a substantially livier habitus after treatment.
Vermedal H, Hellings IR, Fretheim-Kelly ZL, Fintl C, Olsen HMB, Strand E.Exercise-related upper respiratory tract (URT) disorders are common in racehorses. Objective assessment of URT mechanics is essential to quantify degrees of obstruction caused by URT disorders identified upon dynamic endoscopy. Objective: To establish reference values for inspiratory and expiratory tracheal pressures (cmHO) during high-speed treadmill endoscopy in harness racehorses with clinically normal URTs. Methods: Prospective observational study. Methods: Tracheal pressures were measured in harness racehorses (Standardbreds and Norwegian-Swedish coldblooded trotters) in which no URT abno...
Andersson-Eklund L, Andersson L, Sandberg K.A study was carried out to attempt to explain the basis of the association between the Es locus and starting proportion in Swedish Trotters which had been observed previously. The effect of Es genotype on starting proportion has diminished among horses born in the late 1970s. There are indications that the incidence of leg lesions varies between Es genotypes. If this is true, it is possible that the decreasing effect of the Es locus on starting proportion is due to the environmental changes which have been made at race tracks in the early 1980s to reduce the strain on the legs of the trotters....
Varewyck H, Bouquet Y, Lazary S, Guérin G, Van de Weghe A, Van Zeveren A.158 Belgian Saddlebreds, 130 Belgian Trotters, 108 Belgian Draft horses and 92 Shetland ponies have been typed for serologically defined antigens at the ELA and ELY systems. Gene frequencies were estimated in each breed for the internationally established ELA, ELY-1 and ELY-2 alleles as well as for locally assigned additional ELA markers and for subtypes of ELA-W3, W9 and W11. The distribution of ELA alleles was in agreement with the expected Hardy-Weinberg equilibrium for the 4 horse breeds described here. Differences in gene frequencies between these main Belgian horse populations were obser...
Kowalik S, Kedzierski W.The effect of interval vs. continuous exercise on plasma leptin and ghrelin concentration in young Standardbred horses was studied. The experiment was conducted on 27 trotters, in the age between 2 and 3 years. They were divided into two groups according to the type of exercise. Blood samples were collected through jugular venipuncture in the following experimental conditions: at rest, immediately after exercise and 30 minutes after the end of the effort. Plasma leptin and ghrelin concentrations were determined using RIA tests. The continuous exercise induced an increase in plasma leptin conce...
Hjertén G, Drevemo S.Compressions in the forelimb of a horse trotting at 3.7 m/s were studied using high-speed cinematography. From a diagram of shortenings it has been concluded that successive loadings in the longitudinal direction from below signify longer impact time and smaller impact forces compared to a theoretically stiff limb. It is proposed that the limb takes up the impact force passively until the onset of loading at the elbow joint, i.e. 30 ms following first contact.
Frietsch G, Weigand E, Prüstel N.The postnatal iron status of 20 trotter foals (7 female, 13 male) was studied by analyzing different parameters in blood drawn on the day of birth, and 14 +/- 2 and 28 +/- 2 days later. Hemoglobin hematocrit, plasma iron and transferrin saturation averaged 14.9 g/dl, 41.8%, 300 micrograms/dl and 59.4%, respectively, on the day of birth. These values were significantly reduced after 14 days, but had approached again the initial levels by day 28. There was a slight, statistically nonsignificant rise in the mean total iron-binding capacity over the four-week period. Iron status was not affecting ...
Wiemer P, Ugahary F.A 6-year-old trotter gelding presented with exercise intolerance and swelling of the left side of the head during exercise and grazing. The complaints were caused by a complete thrombosis of the left jugular vein. In this case report a surgical approach is described in which a bypass was created with a synthetic vascular graft. After treatment the horse was capable of racing at his former level. The patency of the synthetic graft was 11 months.
Moore LV, Zsoldos RR, Licka TF.The present study investigated accelerations of the front and hind hooves of horses comparing two different shoe types. A standard steel shoe, with studs, pins, and in some instances with toe grabs, was compared to a steel shoe covered on the bottom with a layer of polyurethane. Four horses were used; they trotted in hand on an asphalt track at their self-selected speed. The results showed significantly reduced decelerations during the stance phase with the polyurethane-covered shoes (10th percentile median steel -2.77 g, polyurethane -2.46 g; 0.06) and significantly increased decelerations i...
Fazio F, Ferrantelli V, Piccione G, Caola G.In the athletic horse, the efficiency of physical exercise during activity is strictly
connected to several organic–functional characteristics. Among these, cardiovascular
function takes a primary role in determining the quality of performance. The morphological and functional changes occurring in the cardiovascular system during physical
exercise are due to the increasing demand required to deal with intense muscular
activity. Thus, the cardiovascular physiological response to physical exercise is characterized both by transitory modifications and adjustments that are similar in both
t...
Cotrel C, Leleu C, Courouce-Malblanc A.A close relationship between limb and respiratory rhythms has been shown in clinically healthy galloping horses due to mechanical constraints in the thoracic region. This synchronisation leads to a 1/1 ratio between stride frequency (SF) and respiratory frequency (RF) during galloping. Very little is known about locomotor-respiratory coupling (LRC) during fast trot. Objective: To investigate stride and respiratory rates during a standardised exercise test on the track in Standardbred Trotters. Methods: Forty-four French trotters age 2-10 years performed a standardised exercise test consisting ...
Müller C.During the last years, trotting races met with massive criticism, because some of the numerous items of equipment were not to be reconciled with animal protection. In March 1998, the Trotting Association (HVT), as well as the list of authorized equipment. The so-called "positive list" includes harnesses, bridles, bits, auxiliary reins, driving reins, whips and other aids, among other things brushing boots, ear-muffs and head-pole. Especially criticized items are submitted to an assessment according to the present animal protection law. Moreover, possibilities are shown to make positive conditi...
Hyun SH, Ryew CC.The aim of this study was to investigate the motor ability of forelimb according to on- or off-riding during cadences (walk and trot) of horse. Horses and rider selected as subject consisted of total 37 heads of Jeju native horse and 1 female rider. The variables analyzed composed of 1 stride length, 1 step length, elapsed time of stance, elapsed time of swing, elapsed time of 1 step, and forward velocity (x-axis). Two-way analysis of variance of variables was employed for the statistical analysis with the level of significance set at 5% (P<0.05). Trot cadence showed significant difference ...