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Topic:Cell Proliferation

Cell proliferation in horses refers to the process by which cells divide and multiply, contributing to growth, development, and tissue repair. This biological process is fundamental to maintaining normal physiological functions and responding to injuries or diseases. In equine research, cell proliferation is studied to understand its role in various contexts such as wound healing, regenerative medicine, and cancer. Factors influencing cell proliferation in horses include genetic, environmental, and nutritional elements. This page assembles peer-reviewed research studies and scholarly articles that explore the mechanisms, regulation, and implications of cell proliferation in equine health and disease management.
Gastrointestinal neuroendocrinology.
The Veterinary clinics of North America. Equine practice    January 8, 2005   Volume 18, Issue 2 205-217 doi: 10.1016/s0749-0739(02)00008-1
Schneider DA, Sayegh AI.There exists individual enteroendocrine cells spread throughout the gastrointestinal mucosa that release specific peptide, as well as nonpeptide, hormones to have various endocrine action on target cells bearing cell surface receptors selectively sensitive to these regulatory substances. Following receptor activation, a series of events is set into motion that serves to transduce the information imparted to the target cell. Such transduction mechanisms are numerous, and may be excitatory or inhibitory to the cell depending upon which G-protein subunits the receptor is coupled.
The proliferation inhibitory proteins p27(Kip1) and retinoblastoma are involved in the control of equine lymphocyte proliferation.
Veterinary immunology and immunopathology    November 16, 2004   Volume 102, Issue 4 363-377 doi: 10.1016/j.vetimm.2004.07.001
Flaminio MJ, Yen A, Antczak DF.Observations in early equine pregnancy clearly reveal maternal immune recognition of and response to the presence of the conceptus. Nevertheless, both maternal cellular and humoral responses appear ineffective in destroying the developing placenta and fetus in early pregnancy. Our previous studies had shown that the pre-conditioned medium generated from the culture of equine invasive trophoblast inhibited mitogen-induced lymphocyte proliferation and the expression of cytokine messenger RNA in vitro. Those findings also suggested that lymphocytes might have been halted in the G0/G1 phase of the...
Tenocyte response to cyclical strain and transforming growth factor beta is dependent upon age and site of origin.
Biorheology    October 13, 2004   Volume 41, Issue 5 613-628 
Goodman SA, May SA, Heinegård D, Smith RK.The effect of strain and transforming growth factor beta on equine tendon fibroblasts (tenocytes) was assessed in vitro. Tenocytes were isolated from flexor and extensor tendons of horses from foetal to 10 years of age. These cells were cultured until confluent on collagen-coated silicone dishes. Cyclic biaxial strain of 9+/-1% was applied at 0.5 Hz for 24 hours with or without added TGFbeta1 or 3 (10 ng/ml). Proliferation and synthetic responses were dependent on the tendon of origin. Neither strain nor TGFbeta caused flexor tenocytes to proliferate significantly, while strain alone did proli...
Topological assignment of the N-terminal extension of plasma gelsolin to the gelsolin surface.
The Biochemical journal    September 21, 2004   Volume 385, Issue Pt 3 659-665 doi: 10.1042/BJ20040875
Fock U, Jockusch BM, Schubert WD, Hinssen H.The actin-binding protein gelsolin is highly conserved in vertebrates and exists in two isoforms, a cytoplasmic and an extracellular variant, generated by alternative splicing. In mammals, these isoforms differ only by an N-terminal extension in plasma gelsolin, a short sequence of up to 25 amino acids. Cells and tissues may contain both variants, as plasma gelsolin is secreted by many cell types. The tertiary structure of equine plasma gelsolin has been elucidated, but without any information on the N-terminal extension. In this paper, we present topographical data on the N-terminal extension...
Engineered articular cartilage: influence of the scaffold on cell phenotype and proliferation.
Journal of materials science. Materials in medicine    September 7, 2004   Volume 14, Issue 8 713-716 doi: 10.1023/a:1024915817061
Gigante A, Bevilacqua C, Cappella M, Manzotti S, Greco F.Articular cartilage defects do not heal. Biodegradable scaffolds have been studied for cartilage engineering in order to implant autologous chondrocytes and help cartilage repair. We tested some new collagen matrices differing in collagen type, origin, structure and methods of extraction and purification, and compared the behavior of human chondrocytes cultured on them. Human chondrocytes were grown for three weeks on four different equine type I collagen matrices, one type I, III porcine collagen matrix and one porcine type II collagen matrix. After 21 days, samples were subjected to histoche...
Proliferative aspects of airway smooth muscle.
The Journal of allergy and clinical immunology    August 17, 2004   Volume 114, Issue 2 Suppl S2-S17 doi: 10.1016/j.jaci.2004.04.039
Hirst SJ, Martin JG, Bonacci JV, Chan V, Fixman ED, Hamid QA, Herszberg B, Lavoie JP, McVicker CG, Moir LM, Nguyen TT, Peng Q, Ramos-Barbón D....Increased airway smooth muscle (ASM) mass is perhaps the most important component of the airway wall remodeling process in asthma. Known mediators of ASM proliferation in cell culture models fall into 2 categories: those that activate receptors with intrinsic receptor tyrosine kinase activity and those that have their effects through receptors linked to heterotrimeric guanosine triphosphate-binding proteins. The major candidate signaling pathways activated by ASM mitogens are those dependent on extracellular signal-regulated kinase and phosphoinositide 3'-kinase. Increases in ASM mass may also...
Aquaporin water channels AQP1 and AQP3, are expressed in equine articular chondrocytes.
Veterinary journal (London, England : 1997)    August 11, 2004   Volume 168, Issue 2 143-150 doi: 10.1016/j.tvjl.2003.08.001
Mobasheri A, Trujillo E, Bell S, Carter SD, Clegg PD, Martín-Vasallo P, Marples D.Chondrocytes exist in an unusual and highly variable ionic and osmotic environment in the extracellular matrix of articular cartilage. Alterations to the ionic and osmotic environment of chondrocytes influence the volume and ionic content of the cells, which, in turn, modifies the rate at which extracellular matrix macromolecules are synthesized and degraded. Thus, regulation of the water and solute content of chondrocytes will profoundly affect their anabolic and catabolic functions. The water content of cells is effectively influenced by the abundance of aquaporin (AQP) water channels. Recen...
Cell-mediated immune responses in horses with equine protozoal myeloencephalitis.
The Journal of parasitology    May 29, 2004   Volume 90, Issue 2 428-430 doi: 10.1645/GE-3289RN
Spencer JA, Ellison SE, Guarino AJ, Blagburn BL.Equine protozoal myeloencephalitis (EPM) is a neurologic syndrome seen in horses from the Americas and is mainly caused by Sarcocystis neurona. Cell-mediated immune responses to mitogens have been shown to be reduced in horses with EPM, although it is not known whether the parasite causes this immunosuppression or if the immunosuppression is required for disease manifestation. Recently, a 29-kDa surface antigen from S. neurona merozoites was identified as being highly immunodominant on Western blot. This antigen has been sequenced and cloned, and the expressed protein has been named SnSAG1. Is...
Germ cell apoptosis in the testes of normal stallions.
Theriogenology    May 26, 2004   Volume 62, Issue 1-2 283-297 doi: 10.1016/j.theriogenology.2003.10.022
Heninger NL, Staub C, Blanchard TL, Johnson L, Varner DD, Forrest DW.Apoptosis in testicular germ cells has been demonstrated in many mammalian species. However, little is known about the stallion (Equus caballus) and rates of apoptosis during spermatogenesis. Morphological and biochemical features of apoptosis reported in other species were used to confirm that the TdT-mediated dUTP Nick end labeling (TUNEL) assay is an acceptable method for identification and quantification of apoptotic germ cells in histological tissue sections from stallion testis. Seminiferous tubules from eight stallions with normal testis size and semen quality were evaluated according t...
Hoof wall wound repair.
Equine veterinary journal    May 19, 2004   Volume 36, Issue 3 210-215 doi: 10.2746/0425164044877189
Pollitt CC, Daradka M.Surgical stripping of the hoof wall results in a wound that heals remarkabley well. In contrast, lamellae recovering from laminitis are often deformed. Investigating lamellar wound healing may aid understanding of laminitis. Objective: To document temporal changes in the lamellar basement membrane (BM), dermis and epidermis after surgery. Methods: Wall strips were made in the dorsal hoof wall midline of 6 mature horses. Immunohistochemistry was used to document changes in the basement membrane (BM) and detect proliferation of epidermal cells in lamellar tissues harvested at intervals. A confor...
Epidermal cell proliferation in the equine hoof wall.
Equine veterinary journal    May 19, 2004   Volume 36, Issue 3 236-241 doi: 10.2746/0425164044877198
Daradka M, Pollitt CC.Current theories explaining how the hoof wall 'grows' and moves past the stationary distal phalanx are speculative and based on incomplete evidence. Movement in the lamellar region could occur by cell proliferation or an enzyme-based remodelling process. Since laminitis pathogenesis appears to involve increased transcription and activation of enzymes normally involved in tissue remodelling, it is important to know precisely which process dominates the lamellar region of the hoof.. Objective: To investigate epidermal cell proliferation in the equine hoof wall and calculate a proliferative index...
The effect of enrofloxacin on cell proliferation and proteoglycans in horse tendon cells.
Cell biology and toxicology    May 4, 2004   Volume 20, Issue 1 41-54 doi: 10.1023/b:cbto.0000021154.01035.f9
Yoon JH, Brooks RL, Khan A, Pan H, Bryan J, Zhang J, Budsberg SC, Mueller PO, Halper J.Fluoroquinolone antibiotics have been used widely in humans and domestic animals, including horses, because of their broad-spectrum bactericidal activity, and relative safety. The use of fluoroquinolones, however, is not without risk. Tendonitis and spontaneous tendon rupture have been reported in people during or following therapy with fluoroquinolones. We have studied the effects of enrofloxacin, a fluoroquinolone antibiotic used commonly in domestic animals, on tendon cell cultures established from equine superficial digital flexor tendons. Effects on cell proliferation and morphology were ...
Biochemical and functional assessment of equine lymphocyte phosphodiesterases and protein kinase C.
Veterinary immunology and immunopathology    March 11, 2004   Volume 98, Issue 3-4 153-165 doi: 10.1016/j.vetimm.2003.12.001
Rickards KJ, Page CP, Hamblin AS, Goode NT, Cunningham FM.Lymphocytes play an important role in allergic inflammation and have been implicated in the pathogenesis of equine allergic skin and respiratory disease. Targeting intracellular signalling pathways in human lymphocytes has demonstrated a role for both phosphodiesterase and protein kinase C in cell activation. The aim of this study was to measure total cyclic nucleotide hydrolysing phosphodiesterase activity and to identify the phosphodiesterase and protein kinase C isoenzymes present in equine lymphocytes. The functional significance of these isoenzymes was then investigated by examining their...
Effects of insulin-like growth factor-II on the mitogenic and metabolic activities of equine articular cartilage with and without interleukin 1-beta.
American journal of veterinary research    February 21, 2004   Volume 65, Issue 2 238-244 doi: 10.2460/ajvr.2004.65.238
Davenport-Goodall CL, Boston RC, Richardson DW.To investigate the effects of insulin-like growth factor-II (IGF-II) on DNA and glycosaminoglycan (GAG) synthesis and the expression of matrix-related genes in equine articular cartilage explants and chondrocytes, respectively, with and without interleukin 1-beta (IL1-beta). Methods: Articular cartilage from 12 adult horses. Methods: Articular cartilage was incubated in standard media with and without equine IL1-beta (10 ng/mL) containing various concentrations of IGF-II for 72 hours. Synthesis of DNA and GAG was determined by incorporation of thymidine labeled with radioactive hydrogen (3H) a...
Adhesion and proliferation of human dermal fibroblasts on collagen matrix.
Journal of biomaterials applications    February 12, 2004   Volume 18, Issue 3 209-222 doi: 10.1177/0885328204039692
Croce MA, Silvestri C, Guerra D, Carnevali E, Boraldi F, Tiozzo R, Parma B.The purpose of this study was to evaluate adhesion and growth of human dermal fibroblasts on a 0.150 mm-thick matrix of reconstituted collagen isolated from horse tendon. Collagen was extracted and polymerized according to the standard procedures (Opocrin, Corlo, Modena, Italy). By light microscopy, the bottom surface of the matrix appeared linear and compact, whereas the superficial one was indented and less homogeneous. By scanning electron microscopy, the collagen fibrils had different diameters and the great majority of them was oriented parallel to the surface of the gel. By transmission ...
Cross-species reactivity of seven monoclonal antibodies with equine lymphocytes by flow cytometry.
Veterinary research    January 30, 2004   Volume 34, Issue 6 791-801 doi: 10.1051/vetres:2003033
Mérant C, Bonnefont C, Desbos A, Greenland T, Cadoré JL, Monier JC.The recognition of equine lymphocyte antigens by monoclonal antibodies (mAbs) directed against human CD11a, CD18, CD21, CD23, CD29 and DR, as well as mouse CD23 was studied by flow cytometry. Unlike anti-CD11a, -CD21, -CD23 and DR mAbs, anti-CD18 and CD29 mAbs labelled the same percentage of horse peripheral blood lymphocytes (PBL) as human PBL. Double-staining with anti-horse immunoglobulin antibodies showed that anti-CD21 and -CD23 mAbs are mainly bound to peripheral blood B lymphocytes. The seven mAbs were also tested on the lymph node and thymus cells. The molecular targets of anti-CD11a, ...
Generation and characterisation of an equine macrophage cell line (e-CAS cells) derived from equine bone marrow cells.
Veterinary immunology and immunopathology    January 1, 2004   Volume 97, Issue 1-2 65-76 doi: 10.1016/j.vetimm.2003.08.012
Werners AH, Bull S, Fink-Gremmels J, Bryant CE.Macrophages play a pivotal role in the pathophysiology of many diseases by mediating the host immune response to infections and intoxications. The species-specific activation of macrophages and the differential response in cytokine production impedes the extrapolation of results between species. Therefore, the aim of this study was to isolate and immortalise macrophages from equine bone marrow (BM) cells in order to study equine-specific signalling pathways. The isolated BM-derived macrophages (referred to as e-CAS cells) showed proliferation kinetics similar to that of standardised cell lines...
Comparison of perioperative versus postoperative intratumoral administration of cisplatin for treatment of cutaneous sarcoids and squamous cell carcinomas in horses.
Journal of the American Veterinary Medical Association    October 22, 2003   Volume 215, Issue 11 1655-1660 
Théon AP, Pascoe JR, Galuppo LD, Fisher PE, Griffey SM, Madigan JE.To determine the benefits of reducing the interval between surgical cytoreduction and intratumoral administration of cisplatin. Methods: Randomized clinical study. Methods: 70 horses with 89 incompletely resected T2- and T3-stage sarcoids (n = 64) and squamous cell carcinomas (25). Methods: Horses were given 4 intratumoral treatments of cisplatin at 2-week intervals. The first treatment was given at the time of, or immediately after, surgical resection for horses treated in accordance with the perioperative protocol (group 1). Horses in group 2 were treated with cisplatin after the skin healed...
Cloning, expression and biological activity of equine interleukin (IL)-5.
Veterinary immunology and immunopathology    September 13, 2003   Volume 95, Issue 1-2 63-72 doi: 10.1016/s0165-2427(03)00100-4
Cunningham FM, Vandergrifft E, Bailey SR, Sepulveda MF, Goode NT, Horohov DW.The cytokine, interleukin (IL)-5 stimulates eosinophil differentiation, activation and survival and can prime these cells, increasing the response to other mediators. In view of its many effects on eosinophils, IL-5 has been implicated in the pathogenesis of allergic disease in man. Here we report the cloning of equine IL-5 and expression of the recombinant protein by transfection of Chinese hamster ovary (CHO) cells. The cloned cDNA sequence consisted of 405 nucleotides and encoded a protein of 135 amino acids. There is >85% identity with feline, bovine, ovine, canine, and human IL-5 sequence...
Effects of growth factors (EGF, PDGF-BB and TGF-beta 1) on cultured equine epithelial cells and keratocytes: implications for wound healing.
Veterinary ophthalmology    September 3, 2003   Volume 6, Issue 3 211-217 doi: 10.1046/j.1463-5224.2003.00296.x
Haber M, Cao Z, Panjwani N, Bedenice D, Li WW, Provost PJ.The physiologic mechanisms involving growth factors, including PDGF-BB, EGF, and TGF-beta 1, as potent mediators of fibroblasts and epithelial cells in corneal wound healing remain unknown. The goal of this study was to determine culture methods for equine epithelial cells and keratocytes and to investigate how exogenous growth factors influence proliferation of both cell types. Methods: Cell cultures were established from healthy corneas harvested from horses immediately following euthanasia and maintained using standard tissue culture protocols. To determine the effects of PDGF-BB, EGF, TGF-...
Activation of equine nuclear transfer oocytes: methods and timing of treatment in relation to nuclear remodeling.
Biology of reproduction    September 3, 2003   Volume 70, Issue 1 46-53 doi: 10.1095/biolreprod.103.018200
Choi YH, Love LB, Westhusin ME, Hinrichs K.Early development of embryos produced by transfer of equine nuclei to bovine cytoplasts is superior to that of intraspecies equine nuclear transfer embryos. This may be related to differences in chromatin remodeling or efficiency of activation between the two oocyte types. The pattern of donor nucleus remodeling was examined in equine-equine and equine-bovine reconstructed oocytes. Chromosome condensation occurred in equine cytoplasts by 2 h but was not seen in bovine cytoplasts until 4 h. We investigated the effect of activation of equine-equine reconstructed oocytes at <30 min or at 2 h a...
Immunologic analysis of blood samples obtained from horses and stored for twenty-four hours.
American journal of veterinary research    August 21, 2003   Volume 64, Issue 8 1003-1009 doi: 10.2460/ajvr.2003.64.1003
Witonsky S, Gogal RM, Buechner-Maxwell V, Ahmed SA.To determine whether immune function can be accurately assessed in blood samples obtained from horses and refrigerated overnight and whether a nonradioactive lymphocyte proliferation assay can be used to evaluate samples obtained from horses. Methods: 224 blood samples from 28 clinically normal adult horses. Methods: Heparinized blood samples were collected. Each sample was divided into 2 equal aliquots. One aliquot was refrigerated overnight to simulate overnight shipping of blood samples, and the other aliquot was evaluated on the day of blood collection. Lymphocytes were isolated and enumer...
Equine articular cartilage chondrocytes: opening the black box.
Equine veterinary journal    July 24, 2003   Volume 35, Issue 5 425-428 doi: 10.2746/042516403775600442
Hall AC, Bush PG, Davidson ME, Kempson SA.No abstract available
Dynamic compressive strain inhibits nitric oxide synthesis by equine chondrocytes isolated from different areas of the cartilage surface.
Equine veterinary journal    July 24, 2003   Volume 35, Issue 5 451-456 doi: 10.2746/042516403775600532
Wiseman M, Henson F, Lee DA, Bader DL.Chondrocytes within articular cartilage respond to the mechanical stresses associated with normal joint loading via a series of signalling pathways. Specific biomolecules, such as nitric oxide (NO), have been implicated in these mechanotransduction processes. It has been shown that the synthesis of NO can be inhibited by dynamic compressive strain of chondrocytes in vitro which, in turn, leads to an up-regulation of specific metabolic parameters. Objective: Chondrocytes isolated from different joint locations and seeded in agarose constructs respond in a distinct manner to the application of d...
Clostridium perfringens alpha-toxin-induced hemolysis of horse erythrocytes is dependent on Ca2+ uptake.
Biochimica et biophysica acta    July 2, 2003   Volume 1613, Issue 1-2 79-86 doi: 10.1016/s0005-2736(03)00140-8
Ochi S, Oda M, Nagahama M, Sakurai J.Clostridium perfringens alpha-toxin is able to lyse various erythrocytes. Exposure of horse erythrocytes to alpha-toxin simultaneously induced hot-cold hemolysis and stimulated production of diacylglycerol and phosphorylcholine. When A23187-treated erythrocytes were treated with the toxin, these events were dependent on the concentration of extracellular Ca2+ . Incubation with the toxin of BAPTA-AM-treated horse erythrocytes caused no hemolysis or production of phosphorylcholine, but that of the BAPTA-treated erythrocytes did. When Quin 2-AM-treated erythrocytes were incubated with the toxin i...
[The occurrence of hemodynamic effective elements in the intestinal blood vessel system].
Anatomia, histologia, embryologia    June 12, 2003   Volume 32, Issue 2 94-97 doi: 10.1046/j.1439-0264.2003.00463.x
Wille KH, Schnorr B.Arterioles, precapillary sphincters, capillary endothelium, and pericytes probably regulate the blood flow in the intestinal microvascular bed similar to other regions of the body because of their equipment with contractile filaments. Only throttle veins with their arrangement of pools and their characteristics probably exert influence on the hemodynamic qualities of the blood flow in the intestinal mucosa.
Equine telomeres and telomerase in cellular immortalisation and ageing.
Mechanisms of ageing and development    June 5, 2003   Volume 124, Issue 6 759-764 doi: 10.1016/s0047-6374(03)00104-0
Argyle D, Ellsmore V, Gault EA, Munro AF, Nasir L.To determine the role of telomeres in cellular ageing in equids, we analysed telomere lengths in peripheral blood derived DNA samples from a panel of donkeys (Equus asinus) ranging from 2 to 30 years of age. The average telomere lengths ranged from 7 to 21 kbp and a statistically significant inverse correlation between telomere lengths and donor age was demonstrated. Similarly, telomere lengths in primary fibroblasts isolated from a horse (Equus equus) demonstrated telomeric loss with in vitro ageing when cultured to senescence. We extended this study to evaluate activity of the enzyme telomer...
A mule cloned from fetal cells by nuclear transfer.
Science (New York, N.Y.)    May 29, 2003   Volume 301, Issue 5636 1063 doi: 10.1126/science.1086743
Woods GL, White KL, Vanderwall DK, Li GP, Aston KI, Bunch TD, Meerdo LN, Pate BJ.No abstract available
Substance P induces activation, adherence and migration of equine eosinophils.
Journal of veterinary pharmacology and therapeutics    April 2, 2003   Volume 26, Issue 2 131-138 doi: 10.1046/j.1365-2885.2003.00453.x
Foster AP, Cunningham FM.The tachykinin, substance P (SP), affects eosinophil function by direct and indirect mechanisms and has been shown to cause equine eosinophils to adhere to vascular endothelium and to release cytokines that increase cell adherence. The aim of this study was to determine whether SP could act directly on equine eosinophils in vitro. Eosinophil activation was also compared in cells from normal ponies and those with insect hypersensitivity as SP may be released in the skin of hypersensitive animals. SP caused equine eosinophils to adhere, migrate and produce superoxide, although high concentration...
Analysis of atresia in equine follicles using histology, fresh granulosa cell morphology and detection of DNA fragmentation.
Reproduction (Cambridge, England)    March 4, 2003   Volume 125, Issue 3 417-423 
Pedersen HG, Watson ED, Telfer EE.Follicular atresia has been examined previously by various biochemical and histological methods. The aim of this study was to compare, for the first time, detection of granulosa cell apoptosis by biochemical DNA analysis and microscopic examination of fresh granulosa cell morphology with the established method of detecting atresia by histology in equine follicles. DNA extracted from granulosa cells was examined by staining with ethidium bromide and end-labelling with [(32)P]dideoxy-ATP, which labels the free 3'-end of DNA fragments. In 25 of 26 follicles (96%) there was agreement between end-l...
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