Equine Studies encompasses the scientific exploration and analysis of various aspects related to horses, including their physiology, behavior, genetics, nutrition, and management. This interdisciplinary field integrates knowledge from veterinary medicine, animal science, and equine management to enhance understanding of horse health and welfare. Topics within equine studies often include the study of equine anatomy, disease prevention, breeding practices, and performance optimization. Researchers and scholars contribute to this field by conducting experiments, field studies, and reviews that provide insights into improving equine care and management practices. This page compiles peer-reviewed research studies and scholarly articles that investigate diverse areas within equine studies, offering comprehensive insights into the complexities of horse biology and management.
Rovere G, Ducro BJ, van Arendonk JA, Norberg E, Madsen P.Most Warmblood horse studbooks aim to improve the performance in dressage and show jumping. The Dutch Royal Warmblood Studbook (KWPN) includes the highest score achieved in competition by a horse to evaluate its genetic ability of performance. However, the records collected during competition are associated with some aspects that might affect the quality of the genetic evaluation based on these records. These aspects include the influence of rider, censoring and preselection of the data. The aim of this study was to quantify the impact of rider effect, censoring and preselection on the genetic...
Bocian K, Strzelec K, Janczarek I, Jabłecki Z, Kolstrung R.This paper analyzes changes in the length of coat on selected body areas in horses and ponies kept under different husbandry (stable) conditions during the winter-spring period. The study included 12 Małpolski geldings and 12 geldings of Felin ponies aged 10-15 years. Horses were kept in two stables (six horses and six ponies in each stable). The type of performance, husbandry conditions and feeding of the studied animals were comparable. As of December 1, samples of hair coat from the scapula, sternum, back and abdomen areas of both body sides were collected seven times. The lengths of 20 ra...
François L, Jäderkvist Fegraeus K, Eriksson S, Andersson LS, Tesfayonas YG, Viluma A, Imsland F, Buys N, Mikko S, Lindgren G, Velie BD.Many genes are known to have an influence on conformation and performance traits; however, the role of one gene, Myostatin (MSTN), has been highlighted in recent studies on horses. Myostatin acts as a repressor in the development and regulation of differentiation and proliferative growth of skeletal muscle. Several studies have examined the link between MSTN, conformation, and performance in racing breeds, but no studies have investigated the relationship in Icelandic horses. Icelandic horses, a highly unique breed, are known both for their robust and compact conformation as well as their addi...
Stefanetti V, Marenzoni ML, Passamonti F, Cappelli K, Garcia-Etxebarria K, Coletti M, Capomaccio S.Endogenous retroviruses (ERVs) are proviral phases of exogenous retroviruses that have co-evolved with vertebrate genomes for millions of years. Previous studies have identified the envelope (env) protein genes of retroviral origin preferentially expressed in the placenta which suggests a role in placentation based on their membrane fusogenic capacity and therefore they have been named syncytins. Until now, all the characterized syncytins have been associated with three invasive placentation types: the endotheliochorial (Carnivora), the synepitheliochorial (Ruminantia), and the hemochorial pla...
Christensen BW, Danielson JA.Recent publications have revealed inadequacies in the veterinary training of future equine practitioners. To help address this problem, a 2-week Equine Clinical Skills course was designed and implemented to provide fourth-year veterinary students with opportunities to have hands-on experience with common equine clinical skills using live animals and cadavers. Alumni and employers of alumni were surveyed to determine whether or not students participating in the course were more competent performing clinical skills during their first year post-graduation than those who had not participated in th...
Graham R, McManus P.Changing social values and new technologies have contributed to increasing media attention and debate about the acceptable use of animals in sport. This paper focuses on the use of the whip in thoroughbred horse racing. Those who defend its use argue it is a necessary tool needed for safety, correction and encouragement, and that it does not cause the horse any pain. For those who oppose its use, it is an instrument of cruelty. Media framing is employed to unpack the discourses played out in print and social media in the UK (2011) and Australia (2009) during key periods of the whip debate foll...
Lubow BC, Ransom JI.Reliably estimating wildlife abundance is fundamental to effective management. Aerial surveys are one of the only spatially robust tools for estimating large mammal populations, but statistical sampling methods are required to address detection biases that affect accuracy and precision of the estimates. Although various methods for correcting aerial survey bias are employed on large mammal species around the world, these have rarely been rigorously validated. Several populations of feral horses (Equus caballus) in the western United States have been intensively studied, resulting in identifica...
Staiger EA, Abri MA, Silva CA, Brooks SA.Following domestication, man selected the horse primarily for the purpose of transportation rather than consumption; this selective strategy created divergent traits for locomotion. At intermediate speeds, beyond the flat walk, the horse can perform a range of diagonal and lateral 2-beat or 4-beat gait patterns. The Tennessee Walking Horse (TWH) is the only U.S. breed able to perform an even-timed 4-beat gait (the "running-walk") at intermediate speeds; however, within the breed, there is remaining variation in gait type. To investigate the contribution of genetics to this unique trait, blood ...
Fenner K, Yoon S, White P, Starling M, McGreevy P.Restrictive nosebands are common in equestrian sport. This is concerning, as recent evidence suggests that very tight nosebands can cause a physiological stress response, and may compromise welfare. The objective of the current study was to investigate relationships that noseband tightness has with oral behavior and with physiological changes that indicate a stress response, such as increases in eye temperature (measured with infrared thermography) and heart rate and decreases in heart rate variability (HRV). Horses (n = 12) wearing a double bridle and crank noseband, as is common in dressage ...
Younes M, Robert C, Barrey E, Cottin F.Although cardiac recovery is an important criterion for ranking horses in endurance competitions, heart rate variability (HRV) has hardly ever been studied in the context of this equestrian discipline. In the present study, we sought to determine whether HRV is affected by parameters such as age, exercise duration and test site. Accordingly, HRV might be used to select endurance horses with the fastest cardiac recovery. The main objective of the present study was to determine the effects of age, exercise duration, and test site on HRV variables at rest and during exercise and recovery in young...
Harlow BE, Lawrence LM, Hayes SH, Crum A, Flythe MD.Starch from corn is less susceptible to equine small intestinal digestion than starch from oats, and starch that reaches the hindgut can be utilized by the microbiota. The objective of the current study was to examine the effects of starch source on equine fecal microbiota. Thirty horses were assigned to treatments: control (hay only), HC (high corn), HO (high oats), LC (low corn), LO (low oats), and LW (low pelleted wheat middlings). Horses received an all-forage diet (2 wk; d -14 to d -1) before the treatment diets (2 wk; d 1 to 14). Starch was introduced gradually so that horses received 50...
Yarnell K, Purcell RS, Walker SL.Adrenal activity can be assessed in the equine species by analysis of feces for corticosterone metabolites. During a potentially aversive situation, corticotrophin releasing hormone (CRH) is released from the hypothalamus in the brain. This stimulates the release of adrenocorticotrophic hormone (ACTH) from the pituitary gland, which in turn stimulates release of glucocorticoids from the adrenal gland. In horses the glucocorticoid corticosterone is responsible for several adaptations needed to support equine flight behaviour and subsequent removal from the aversive situation. Corticosterone met...
Hinrichs K.A research study is a product of not only a question and its pursuit but also the people, places, and facilities available at the time. My work in equine assisted reproduction has progressed from embryo transfer to oocyte maturation, oocyte transfer, intracytoplasmic sperm injection, embryo biopsy, embryo vitrification, and cloning, as a result of collaborations with an array of remarkable people. This is a summary of some of the stories behind the studies.
Pedersen PJ, Karlsson M, Flethøj M, Trachsel DS, Kanters JK, Klaerke DA, Buhl R.Quantitative measurements of cardiac repolarization, defined as the electrocardiographic QT interval, have important diagnostic implications in humans, as irregularities can trigger potentially fatal ventricular tachyarrhythmia. In both humans and horses, cardiac repolarization is influenced to some extent by heart rate, age, body weight (BW), sex, autonomic tone, and environment. In horses, there is substantial inter-breed variation in size and training, and the aims of this study were therefore to determine the best model describing the QT to RR relationship in breeds of various athletic hor...
The climatic and cultural diversity of the Italian Peninsula triggered, over time, the development of a great variety of horse breeds, whose origin and history are still unclear. To clarify this issue, analyses on phenotypic traits and genealogical data were recently coupled with molecular screening. Methods: To provide a comprehensive overview of the horse genetic variability in Italy, we produced and phylogenetically analyzed 407 mitochondrial DNA (mtDNA) control-region sequences from ten of the most important Italian riding horse and pony breeds: Bardigiano, Esperia, Giara, Lipizzan, Maremm...
Grabski M, McNamara J, Roper D, Warren-Smith C, Busschers E.To describe the volume and dimensions of the bicipital bursa and its position in relation to bony and soft tissue structures, in order to develop a novel medial synoviocentesis approach to the bicipital bursa. Methods: Cadaveric study. Methods: Adult equine cadaver limbs (n=19). Methods: Bicipital bursa dimensions, volume, and relationship to bony structures were obtained from positive contrast computed tomography images after distension of the bursa (n=7). Following an intra-bursal injection of polyurethane resin, the bicipital bursa (n=4) was dissected and its relationship to soft tissue str...
Fazio E, Medica P, Trifiletti C, Ferlazzo A.The objective of this study was to compare in detail the total and free iodothyronines' pattern of mares from the first ovulation of the year over an extended period of 12 weeks. A total of 20 mares were used in the study. The mares were classified into two groups: mares mated at the ovulation (n = 10) used as observational group and mares unmated at the ovulation (n = 10) used as control group. Serum total and free triiodothyronine (T3, fT3) and thyroxine (T4, fT4) levels were measured in baseline conditions at the first ovulation of year and once a week until 12 weeks later. For the experi...
Voigt MA, Hiney K, Richardson JC, Waite K, Borron A, Brady CM.The purpose of this study was to gain a better understanding of stock-type horse show competitors' understanding of welfare and level of concern for stock-type show horses' welfare. Data were collected through an online questionnaire that included questions relating to (a) interest and general understanding of horse welfare, (b) welfare concerns of the horse show industry and specifically the stock-type horse show industry, (c) decision-making influences, and (d) level of empathic characteristics. The majority of respondents indicated they agree or strongly agree that physical metrics should b...
Thompson K, Clarkson L.The extension of research into public practice is enhanced by communication and behaviour change strategies that are consistent with consumer needs and perspectives. To gain support for equine research (or to appreciate the perspectives contributing to disagreement), it is necessary to determine how aware consumers are of research, what research means to them, how they perceive its benefits (if at all) and how they engage with (or resist) it. Because of a surprising dearth of research evaluating consumer perceptions of research in any sector, our aim was to identify the perceived outputs and b...
Clayton HM.The central body axis or core is a key component in controlling body posture and providing a stable platform for limb movements and generation of locomotor forces. Persistent dysfunction of the deep stabilizing muscles seems to be common in horses indicating a need for core training exercises to restore normal function. Core training should be performed throughout the horse's athletic career to maintain a healthy back and used therapeutically when back pain is identified. This article reviews the structure and function of the equine thoracolumbar spine with special reference to function, dysfu...
Brubaker L, Udell MA.Horses (Equus caballus) have a rich history in their relationship with humans. Across different cultures and eras they have been utilized for work, show, cultural rituals, consumption, therapy, and companionship and continue to serve in many of these roles today. As one of the most commonly trained domestic animals, understanding how horses learn and how their relationship with humans and other horses impacts their ability to learn has implications for horse welfare, training, husbandry and management. Given that unlike dogs and cats, domesticated horses have evolved from prey animals, the hor...
Gill Harris reports from this year's National Equine Forum where a key theme was the importance of collaboration and effective communication in achieving the best outcomes for the health and welfare of the horse and the future of equestrianism in the UK.
Negro Rama S, Valera M, Membrillo A, Gómez MD, Solé M, Menendez-Buxadera A, Anaya G, Molina A.The association of five candidate genes with sporting performance in young and adult Spanish Trotter horses (STHs) was performed according to a previous selection based on quantitative analysis of the trait time per kilometre (TPK). A total of 334 516 records of TPK from 5958 STHs were used to estimate the estimated breeding values (EBVs) at different age groups (young and adults horses) throughout the range of distances (1600-2700 m) using a bicharacter random regression model. The heritability estimated by distance ranged from 0.16 to 0.40, with a different range for the two age groups. Co...
Mochizuki M, Hayakawa N, Minowa F, Saito A, Ishioka K, Ueda F, Okubo K, Tazaki H.In this study, iodine and thyroxin (T4) concentrations in the serum of 69 horses were investigated. Higher iodine concentrations were obtained from the horses housed in Chiba Prefecture. In contrast, T4 concentrations of horses at Shizuoka Prefecture were higher than those of horses at Chiba Prefecture. There was a significant correlation (r = 0.643, P < 0.001) between the iodine and T4 concentrations of horses at Saitama and Shizuoka prefectures. Although a significant correlation (r = 0.794, P < 0.001) was also observed in the investigation of all horses at Chiba Prefec...
Sinclair J.Veterinary students with a particular interest in equine medicine and surgery gathered at Nottingham vet school recently to further their knowledge and skills in these areas. Jordan Sinclair, editor of the Journal of the Association of Veterinary Students, reports.
Mach N, Plancade S, Pacholewska A, Lecardonnel J, Rivière J, Moroldo M, Vaiman A, Morgenthaler C, Beinat M, Nevot A, Robert C, Barrey E.The adaptive response to extreme endurance exercise might involve transcriptional and translational regulation by microRNAs (miRNAs). Therefore, the objective of the present study was to perform an integrated analysis of the blood transcriptome and miRNome (using microarrays) in the horse before and after a 160 km endurance competition. A total of 2,453 differentially expressed genes and 167 differentially expressed microRNAs were identified when comparing pre- and post-ride samples. We used a hypergeometric test and its generalization to gain a better understanding of the biological functio...
Gronqvist G, Rogers C, Gee E.Within popular press there has been much coverage of the negative effects associated with firework and horses. The effect of fireworks has been documented in companion animals, yet no studies have investigated the negative effects, or otherwise, of fireworks on horses. This study aims to document horse responses and current management strategies to fireworks via an online survey. Of the total number of horses, 39% (1987/4765) were rated as "anxious", 40% (1816/4765) "very anxious" and only 21% (965/4765) rated as "not anxious" around fireworks. Running (82%, 912/1107) was the most common behav...
Mohite DS, Sheikh CS, Singh S, Kalita J, Williams S, Compston PC.Farriery is a critical component of healthcare services for working equids. However, in India, an informal workforce, lack of structured training facilities and non-implementation of farriery regulations pose challenges for quality farriery. Brooke India, an equine welfare organisation, has undertaken many initiatives aiming to improve farriery services, including technical training and engagement with equid-owning communities. However, this has met with varying success. The study aimed to identify factors that prevent farriers providing quality farriery services. Focus-group discussions were ...
Jones MM, Nuñez CMV.Stress responses can be triggered by several physical and social factors, prompting physiological reactions including increases in glucocorticoid concentrations. In a population of feral horses (Equus caballus) on Shackleford Banks, North Carolina, females previously immunized with the immunocontraceptive agent porcine zona pellucida (PZP) change social groups (bands) more often than unimmunized females, disrupting the social stability within the population. We assessed the effects of increased female group changing behavior (or female turnover) on individual male stress by comparing fecal cor...
Henry S, Briefer S, Richard-Yris MA, Hausberger M.A recent study has shown that gently handling dams in front of their few days old foals may strongly influence the development of human-foal relationships. In the present study, we test whether 6-month-old foals remain sensitive to their dams' influence. The study was performed on 16 foal-mare dyads, with half of the mares receiving positive contacts from the experimenter in presence of their 6-month-old foals (n = 8) whereas the other mares were not handled (n = 8). All foals were tested 15 and 30-35 days later under various conditions (reaction to a motionless human, approach test, saddle-pa...
Hristov P, Yordanov G, Ivanova A, Mitkov I, Sirakova D, Mehandzyiski I, Radoslavov G.It is believed that population structure of mountain horse breeds is preserved from any genetic introgression, because of their geographical isolation and specific semi-wild life style of husbandry. Till date there are no molecular data for the Balkan horses. In this study we try to give information about some autochthonous mountain horse breeds from Bulgaria. A total of 121 horses from three different mountain massifs are presented: Stara Planina (the Balkan mountain), the Rhodopes and Rila-Pirin massif were genotyped according to mitochondrial D-loop region. The results showed huge diversity...
Attenburrow DP.Techniques developed for the detection and simultaneous recording of certain events in the respiratory and limb cycles are presented. The respiratory cycle and limb cycle were found to be in-phase only at the canter and gallop. The significance of any time relationship between events in respiration and in locomotion is discussed.
Davie AJ, Priddle TL, Evans DL.This study was undertaken to invesigate the application of submaximal field tests in assessment of performance. A submaximal field exercise test consisting of 2 bouts of pacing for 1600 m, with 5 min rest or walking between runs was performed on Standardbred racehorses at 2 training centres (A and B). Performance indices were compiled for each horse: number of race starts, number of race wins, number of race placings (1, 2 or 3), and lifetime earnings. Regression analysis was conducted to describe the relationship between plasma lactate concentrations and speed for tests one, 2 and pooled resu...
Hothersall B, Gale EV, Harris P, Nicol CJ.Discrimination learning studies suggest that horses learn more easily using spatial than visible object-specific (OS) cues. However, spatial cues have generally confounded intra-array, distal and/or egocentric spatial information. It is also unclear whether conflicting cues compete for association or are redundantly encoded, and furthermore, the influence of prior experiences or training has not been quantified so far. We examined the effect of cue modality on unweaned foals' performance in a discrimination learning task. After a pilot study confirmed that horses could perform the required OS ...
Easley JT, Hendrickson DA.Equine standing surgery and laparoscopy are becoming increasingly important aspects of equine surgery. Laparoscopic advancements lag behind the human medical field, mainly due to decreased access to appropriate training and instrumentation. It is nearly impossible to cover the topic of equine standing surgery without discussing advances in laparoscopy, because without such advances, equine standing surgery lacks potential for forward progress. Although novel standing techniques continue to be published, the addition of minimally invasive laparoscopic techniques adds an entirely new dimension a...
Attenburrow DP, Goss VA.The phase relationship between the periods of the respiratory and limb cycles is demonstrated in the horse ridden in the field at the canter and gallop. Changes in intra-abdominal pressure, respiratory sounds, periods of ground contact of each foot and volume changes of the rib cage were measured in the normal horse exercised at the walk, trot, canter and gallop in the field. Correlation of these parameters identifies the major mechanical link establishing the coupling of lung ventilation and locomotion in the horse. The force and extent of contraction of the abdominal muscles couples the resp...
Donadeu FX, Ginther OJ.The temporal relationships in the changes in concentrations of follicular fluid factors during follicle selection were characterized in mares. All follicles > or =5 mm were ablated 10 days after ovulation, followed by follicular fluid collection from the three largest follicles (F1, F2, and F3) when F1 of the new wave reached a diameter of 8.0-11.9, 12.0-15.9, 16.0-19.9, 20.0-23.9, 24.0-27.9, or 28.0-31.9 mm (n = 4-8 mares/range). Diameter deviation between F1 and F2 began during the 20.0- to 23.9-mm range, as indicated by a greater difference in diameter between the two follicles at the 24...
Velie BD, Hamilton NA, Wade CM.Different indicators of racing performance are commonly used in the racing industry to assess the genetic superiority of racing Thoroughbreds. However, how well these indicators predict the performance of future progeny or siblings varies depending on the population and circumstances in which the indicators were recorded or achieved. Objective: To identify heritable indicators of racing performance for horses racing in Hong Kong. Methods: Heritability analysis of racing performance traits. Methods: Performance data on the population of Thoroughbreds racing in Hong Kong between 3 September 2000...
Klingel H.The basic social unit in feral horses is the family group consisting of one stallion, one to a few unrelated mares and their foals. Surplus stallions associate in bachelor groups. Stallions are instrumental in bringing mares together in a unit which then persists even without a stallion. The similarity of social organization in populations living in a variety of different habitats indicates that feral horses have reverted to the habits of their wild ancestors, and that domestication has had no influence on this basic behavioural feature.
Muylle S, Simoens P, Lauwers H, Van Loon G.The accuracy of ageing horses by their dentition was assessed by comparing the dental features with the known dates of birth of 212 trotter horses and 189 Belgian draft horses. The horses ranged in age from two to 25 years. In both breeds it was observed that the shedding of the incisors and the appearance of the dental stars were the most reliable features for age determination. In young animals, the dental configuration was similar in both breeds. With increasing age the incisor teeth of draft horses were more liable to abrasion than those of trotter horses. The sequential changes in appeara...
Maśko M, Zdrojkowski L, Domino M, Jasinski T, Gajewski Z.The natural head and neck position (HNP) of horses differs from the position in horse riding when bit is used. The special lunging aids (LAs) are applied in order to modify HNP. Different types of LAs have the potential to affect the work of horse muscles and the superficial thermographic patterns (STPs). The effects of thre LAs on STPs of neck, chest, back, and hindquarters were investigated. Methods: Sixteen leisure horses were lunged with freely moving head (FMH), rubber band (RB), chambon (CH), and triangle side reins (TRs). The thermographic images (n = 896) were analyzed before/after lun...
Drevemo S, Dalin G, Fredricson I, Björne K.The reproducibility of certain gait characteristics was investigated in Standardbred trotters at a speed of 12.0 m/sec. Twenty-two horses were recorded twice on the same day and 4 of them on a third occasion 3 years later. The short-term reproducibility was shown to be very good. In the long-term study it was found that the stride length and duration of swing and stride had increased. This was interpreted as an effect of training.
Devick IF, Leise BS, Rao S, Hendrickson DA.The cause of transient post-operative pain in a subset of horses undergoing laparoscopy is unclear. The objective of this study was to evaluate if residual pneumoperitoneum is associated with transient post-operative pain in mares undergoing ovariectomy. Thirty-eight mares undergoing routine standing laparoscopic ovariectomy were randomly allocated into 2 groups. At the completion of laparoscopy, either the abdominal cavity was actively desufflated or the cannulas were opened to achieve ambient pressure before incisional closure. Assessments were performed for 18 hours after surgery using a va...
Thayer JF, Hahn AW, Pearson MA, Sollers JJ, Johnson PJ, Loch WE.High heart rate variability (HRV) has been associated with more efficient autonomic control, allowing more responsivity and sensitivity to changing environmental demands. Previous results from spectral analysis of interbeat intervals of equine heart rate (HR) indicated increased HRV, reflecting more effective vagal control of the heart. This study focuses on characterizing equine HR and HRV during increasing then decreasing physical demand. A non-invasive ambulatory monitoring system continuously records HR and HRV as horses exercise on a high speed equine treadmill at increasing then decreasi...
Komosa M, Frackowiak H, Purzyc H, Wojnowska M, Gramacki A, Gramacki J.The study included 249 horses belonging to 3 horse breeds. Konik horses, comprising the first group, is an example of a breed similar to the extinct Tarpan. In our study, these horses were taken to be a primitive anatomical model of the horse body. The other groups comprised the Polish Half-bred horse and Thoroughbred horse. The biometric characteristics of the horses were compared based on 24 indices. The aim of the paper was to find a reduced set of indices that can be used to determine group membership of the horses. To do this, we used statistical methods to find the most important indices...
Science (New York, N.Y.)June 24, 1983
Volume 220, Issue 4604 1403-1404 doi: 10.1126/science.220.4604.1403
Berger J.The identities, sexes, and reproductive status of groups of wild horses (Equus caballus) living in the Great Basin Desert of North America were known prior to their deaths on ridgelines. Another group of very young horses died on a quagmire. Snow accumulation or drought was apparently responsible for the mass deaths. These data have implications for reconstructing some aspects of the social structure of fossil mammals on the basis of skewed sex or age ratios in bone assemblages.
Simones P, De Geest JP, Lauwers H.The pectinate ligaments of ten horses, two donkeys, five oxen, five sheep, ten goats, five dogs, five cats, thirty pigs and two rabbits were studied under the stereomicroscope and the scanning electron microscope. In the horse and the donkey, the pectinate ligament was very prominent and was characterized by sturdy interconnected strands and relatively small intertrabecular spaces. The pectinate ligaments of ruminants were composed of shorter strands, separated by relatively larger spaces. Fusion between adjacent strands, resulting in the formation of fenestrated sheets, was regularly observed...
Liu N, Yu W, Guo X, Chen J, Xia D, Yu J, Li D.Lytic polysaccharide monooxygenases (LPMOs) belonging to the auxiliary activity 9 family (AA9) are widely found in aerobic fungi. These enzymes are O-dependent copper oxidoreductases that catalyze the oxidative cleavage of cellulose. However, studies that have investigated AA9 LPMOs of aerobic fungi in the herbivore gut are scare. To date, whether oxidative cleavage of cellulose occurs in the herbivore gut is unknown. Results: We report for the first time experimental evidence that AA9 LPMOs from aerobic thermophilic fungi catalyze the oxidative cleavage of cellulose present in the horse gut t...