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Topic:Locomotion

Locomotion in horses refers to the movement mechanisms that enable horses to walk, trot, canter, and gallop. It involves a complex interplay of the musculoskeletal and nervous systems, allowing for efficient and coordinated movement. Studies on equine locomotion often focus on gait analysis, biomechanics, and the impact of various factors such as conformation, training, and surface conditions on movement. Research in this area contributes to understanding performance, diagnosing lameness, and improving rehabilitation practices. This page compiles peer-reviewed research studies and scholarly articles that explore the biomechanics, gait patterns, and factors influencing locomotion in horses.
The effect of rising and sitting trot on back movements and head-neck position of the horse.
Equine veterinary journal    August 1, 2009   Volume 41, Issue 5 423-427 doi: 10.2746/042516409x371387
De Cocq P, Prinsen H, Springer NC, van Weeren PR, Schreuder M, Muller M, van Leeuwen JL.During trot, the rider can either rise from the saddle during every stride or remain seated. Rising trot is used frequently because it is widely assumed that it decreases the loading of the equine back. This has, however, not been demonstrated in an objective study. Objective: To determine the effects of rising and sitting trot on the movements of the horse. Objective: Sitting trot has more extending effect on the horse's back than rising trot and also results in a higher head and neck position. Methods: Twelve horses and one rider were used. Kinematic data were captured at trot during over gr...
Intensity of activation and timing of deactivation modulate elastic energy storage and release in a pennate muscle and account for gait-specific initiation of limb protraction in the horse.
The Journal of experimental biology    July 21, 2009   Volume 212, Issue Pt 15 2454-2463 doi: 10.1242/jeb.027995
Lichtwark GA, Watson JC, Mavrommatis S, Wilson AM.The equine biceps brachii (biceps) initiates rapid limb protraction through a catapult mechanism. Elastic strain energy is slowly stored in an internal tendon and is then rapidly released to protract the forelimb. The muscle fibres are short, have little scope for length change and can therefore only shorten slowly compared with the speed at which the whole muscle must shorten, which makes them poor candidates for driving rapid limb protraction. We suggest that the muscle fibres in the biceps act to modulate the elastic energy output of the muscle-tendon unit (MTU) to meet the demands of locom...
Trunk deformation in the trotting horse.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 203-206 doi: 10.2746/042516409x393194
Nauwelaerts S, Clayton HM.Estimates of the position of the centres of mass (CM) of body segments are usually extrapolated relative to bony landmarks as determined in cadaver studies. This extrapolation assumes that segments are rigid bodies. Since the trunk represents a large percentage of the total body mass in horses, violation of the rigid body assumption by the trunk segment has important consequences for studying the biomechanics of equine locomotion. Objective: To assess the magnitude of error in CM position due to deformability of the trunk segment and the timing of these errors during the trotting stride. The h...
The effect of different head and neck positions on the caudal back and hindlimb kinematics in the elite dressage horse at trot.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 274-279 doi: 10.2746/042516409x394436
Rhodin M, Gómez Alvarez CB, Byström A, Johnston C, van Weeren PR, Roepstorff L, Weishaupt MA.Dressage involves training of the horse with the head and neck placed in a position defined by the rider. The best position for dressage training is currently under debate among riders and trainers, but there are few scientific data available to confirm or disprove the different views. Objective: To evaluate the kinematic effects of different head and neck positions (HNPs) in elite dressage horses ridden at trot. Methods: Seven high-level dressage horses were subjected to kinetic and kinematic measurements when ridden on a treadmill with the head and neck in 5 different positions. Results: Com...
Kinetics and kinematics of the horse comparing left and right rising trot.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 292-296 doi: 10.2746/042516409x397127
Roepstorff L, Egenvall A, Rhodin M, Byström A, Johnston C, van Weeren PR, Weishaupt M.At rising trot the rider sits alternately down on one diagonal pair of limbs and rises up on the other. The possible effects on asymmetry of locomotion induced by rising trot have rarely been studied. Objective: To demonstrate whether, and if so to what extent, rising trot causes asymmetrical loading in the vertical ground reaction force (VGRF) and/or asymmetrical effects on the locomotion pattern, comparing left and right side. Methods: Seven elite horses were ridden in left and right rising trot on a treadmill, while VGRF and kinematics were measured, with the horses' neck raised, the poll h...
Evaluation of discriminant analysis based on dorsoventral symmetry indices to quantify hindlimb lameness during over ground locomotion in the horse.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 304-308 doi: 10.2746/042516409x397352
Church EE, Walker AM, Wilson AM, Pfau T.Advances in gait analysis techniques have led to assessment tools that can aid in detecting and quantifying lameness; here, bilateral tubera coxae and pelvic movement during over ground locomotion are compared in order to investigate a practical method to assess hindlimb lameness in the horse. Objective: To evaluate which parameters from anatomical landmarks on trunk and proximal hindlimbs are the best indicators of degree and side of hindlimb lameness. Methods: Fifteen horses (age 11-23 years, 6 nonlame and 9 unilaterally hindlimb lame horses 1/10 to 2/10 lame) were fitted with 4 inertial sen...
Back kinematics of healthy trotting horses during treadmill versus over ground locomotion.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 297-300 doi: 10.2746/042516409x397370
Gómez Alvarez CB, Rhodin M, Byström A, Back W, van Weeren PR.Treadmill locomotion is frequently used for training of sport horses, for diagnostic purposes and for research. Identification of the possible biomechanical differences and similarities between the back movement during treadmill (T) and over ground (O) locomotion is essential for the correct interpretation of research results. Objective: To compare the kinematics of the thoracolumbar vertebral column in treadmill and over ground locomotion in healthy horses. Methods: Six sound Dutch Warmblood horses trotted on a T and O during 10 s at their own preferred velocity (mean +/- s.d. 3.6 +/- 0.3 m/s...
The relationship between range of motion of lumbosacral flexion-extension and canter velocity of horses on a treadmill.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 301-303 doi: 10.2746/042516409x397271
Johnson JL, Moore-Colyer M.Research into kinematics of the healthy equine back, has been performed in the walk and trot. This study focuses on back kinematics during canter, over a range of velocities. Flexion extension (FE) movements in canter are greatest in the lumbosacral (LS) region. Previous research has focused on canter velocity of 7 m/s; therefore quantification of LS kinematics at varying velocities is required to understand LS functions in equine locomotion. Objective: Range of flexion-extension movement through the lumbosacral joint increases with increasing velocity. Methods: Six Thoroughbred horses (mean a...
Horizontal moment around the hoof’s centre of pressure during walking in a straight line.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 242-246 doi: 10.2746/042516409x396993
Colborne GR, Heaps LA, Franklin SH.Joint congruity and ligaments restrain the distal limb joints from excessive motion in the transverse and frontal planes, but the magnitudes and direction of the horizontal twisting moments around the hoof's centre of pressure (CoP) that induce these motions are unknown. Objective: To quantify the horizontal moment around the vertical axis through the hoof's CoP at walk, and to determine whether these are symmetric. Methods: Nine sound Thoroughbred horses (mean age 53 years; mean mass 502 kg) were led at walk in a straight line across a Kistler force platform. Five trials were collected for ea...
Skeletal forelimb measurements and hoof spread in relation to asymmetry in the bilateral forelimb of horses.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 238-241 doi: 10.2746/042516409x395561
Wilson GH, McDonald K, O'Connell MJ.Research has highlighted a high frequency of skeletal asymmetries in horses. In addition, research into hoof asymmetries has shown that within a bilateral pair, the hoof with the smaller angle is often subjected to greater loading. There has been limited attention paid to understanding compensatory mechanisms for skeletal asymmetries in the horse; the dynamic structure of the hoof could potentially be acting in a compensatory capacity. Objective: To investigate the relationship between morphometry of forelimb segments and hoof spread and their incidence of asymmetry. Methods: Ten bilateral mea...
Relationship between the forces acting on the horse’s back and the movements of rider and horse while walking on a treadmill.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 285-291 doi: 10.2746/042516409x397136
von Peinen K, Wiestner T, Bogisch S, Roepstorff L, van Weeren PR, Weishaupt MA.The exact relationship between the saddle pressure pattern during one stride cycle and the movements of horse and rider at the walk are poorly understood and have never been investigated in detail. Objective: The movements of rider and horse account for the force distribution pattern under the saddle. Methods: Vertical ground reaction forces (GRF), kinematics of horse and rider as well as saddle forces (FS) were measured synchronously in 7 high level dressage horses while being ridden on an instrumented treadmill at walk. Discrete values of the total saddle forces (FStot) were determined for e...
Effects of a synthetic all-weather waxed track versus a crushed sand track on 3D acceleration of the front hoof in three horses trotting at high speed.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 247-251 doi: 10.2746/042516409x394463
Chateau H, Robin D, Falala S, Pourcelot P, Valette JP, Ravary B, Denoix JM, Crevier-Denoix N.Inadequate track surfaces are believed to be a risk factor in the occurrence of musculoskeletal injuries, but quantification of the shocks and vibrations provoked by hoof impact on different ground surfaces (including new synthetic tracks) has been insufficiently documented in trotters under high-speed training conditions. Objective: To test the reliability and sensitivity of an accelerometric device to discriminate between the biomechanical effects of 2 different tracks at high speed. Methods: Three French Trotters were used and their right front hooves were equipped with one triaxial acceler...
Basic kinematics of the saddle and rider in high-level dressage horses trotting on a treadmill.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 280-284 doi: 10.2746/042516409x394454
Byström A, Rhodin M, von Peinen K, Weishaupt MA, Roepstorff L.A comprehensive kinematic description of rider and saddle movements is not yet present in the scientific literature. Objective: To describe saddle and rider movements in a group of high-level dressage horses and riders. Methods: Seven high-level dressage horses and riders were subjected to kinematic measurements while performing collected trot on a treadmill. For analysis a rigid body model for the saddle and core rider segments, projection angles of the rider's extremities and the neck and trunk of the horse, and distances between markers selected to indicate rider position were used. Results...
Influence of different head-neck positions on vertical ground reaction forces, linear and time parameters in the unridden horse walking and trotting on a treadmill.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 268-273 doi: 10.2746/042516409x397389
Waldern NM, Wiestner T, von Peinen K, Gómez Alvarez CG, Roepstorff L, Johnston C, Meyer H, Weishaupt MA.It is believed that the head-neck position (HNP) has specific effects on the loading pattern of the equine locomotor system, but very few quantitative data are available. Objective: To quantify the effects of 6 different HNPs on forelimb-hindlimb loading and underlying temporal changes. Methods: Vertical ground reaction forces of each limb and interlimb coordination were measured in 7 high level dressage horses walking and trotting on an instrumented treadmill in 6 predetermined HNPs: HNP1--unrestrained; HNP2--elevated neck, bridge of the nose in front of the vertical; HNP3--elevated neck, bri...
Kinetics and kinematics of the passage.
Equine veterinary journal    May 28, 2009   Volume 41, Issue 3 263-267 doi: 10.2746/042516409x397226
Weishaupt MA, Byström A, von Peinen K, Wiestner T, Meyers H, Waldern N, Johnston C, van Weeren R, Roepstorff L.The load acting on the limbs and the load distribution between fore- and hindlimbs while performing specific dressage exercises lack objective assessment. Objective: The greater a horse's level of collection, the more load is shifted to the rear and that during the passage the vertical load on the limbs increases in relation to the accentuated vertical movement of the centre of mass. Methods: Back and limb kinematics, vertical ground reaction force and time parameters of each limb were measured in 6 Grand Prix dressage horses performing on an instrumented treadmill at the trot and the passage....
Motion pattern analysis of gait in horseback riding by means of Principal Component Analysis.
Human movement science    May 13, 2009   Volume 28, Issue 3 394-405 doi: 10.1016/j.humov.2009.04.002
Witte K, Schobesberger H, Peham C.As a consequence of the three interacting systems of horse, saddle, and rider, horseback riding is a very complex movement that is difficult to characterize by a limited number of biomechanical parameters or characteristic curves. Principal Component Analysis (PCA) is a technique for reducing multidimensional datasets to a minimal (i.e., optimally economic) set of dimensions. To apply PCA to horseback riding data, a "pattern vector" composed of the horizontal velocities of a set of body markers was determined. PCA was used to identify the major dynamic constituents of the three natural gaits o...
A comparison of forces acting on the horse’s back and the stability of the rider’s seat in different positions at the trot.
Veterinary journal (London, England : 1997)    May 9, 2009   Volume 184, Issue 1 56-59 doi: 10.1016/j.tvjl.2009.04.007
Peham C, Kotschwar AB, Borkenhagen B, Kuhnke S, Molsner J, Baltacis A.The aim of the study was to compare the stability of the rider as well as the forces acting on a horse's back with different seating positions at the trot (sitting trot, rising trot and two-point seat). The same experienced rider was mounted on 10 sound horses trotting on a treadmill. The kinetic data were recorded with an electronic pressure mat, placed under a well-fitting dressage saddle with no saddle pad. The rider used three different seating positions, each for 20 s. Right forelimb motion was used to synchronise the pressure data with the stride cycles. To determine the rider's stabilit...
An assessment of the pressure distribution exerted by a rider on the back of a horse during hippotherapy.
Human movement science    April 29, 2009   Volume 28, Issue 3 387-393 doi: 10.1016/j.humov.2009.04.001
Janura M, Peham C, Dvorakova T, Elfmark M.Hippotherapy employs locomotion impulses that are emitted from the back of a horse while the horse is walking. These impulses stimulate the rider's postural reflex mechanisms, resulting in training of balance and coordination. The aim of the present study was to assess the changes in magnitude and distribution of the contact pressure between the rider and the horse during a series of hippotherapy lessons. The monitored group, consisting of four healthy women (mean age 22.75 years, mean body weight 59.75 kg, mean height 167.25 cm) without any previous horse riding experience, received five 20 m...
Effects of a stretching regime on stride length and range of motion in equine trot.
Veterinary journal (London, England : 1997)    April 25, 2009   Volume 181, Issue 1 53-55 doi: 10.1016/j.tvjl.2009.03.010
Rose NS, Northrop AJ, Brigden CV, Martin JH.The aim of this study was to quantify the effects of two different 8-week stretching regimes on stride length (SL) and range of motion (ROM) in the equine trot. Eighteen horses were divided into three matched groups: a 6 days/week stretching regime (6DSR), a 3 days/week stretching regime (3DSR) and a control no-stretching regime (NSR). SL and ROM data were collected at weeks 0, 2, 4, 6 and 8 for trot in-hand. Stretching had no significant effect on SL. A number of significant differences were found in joint ROM between treatments in the shoulder, stifle and hock, suggesting some negative biome...
Weighted boots influence performance in show-jumping horses.
Veterinary journal (London, England : 1997)    April 17, 2009   Volume 181, Issue 1 74-76 doi: 10.1016/j.tvjl.2009.03.015
Murphy J.This study investigated the effects of weighted boots on horses (n=6) jumping a 1.25 m oxer fence. The horses had similar training experience and were assigned to two groups of three subjects (groups G1 and G2). All horses performed 10 jumping efforts: G1 horses made attempts 1-5 without boots and 6-10 with boots; G2 made attempts 1-5 with boots and 6-10 without boots. Data were available via sagittal plane S-VHS recordings and t test analyses focussed on limb-placement dimensions. There were no differences among performances of the horses in the horizontal plane, but there were significant di...
SMART: Sensitivity models for animals in response to training.
Veterinary journal (London, England : 1997)    April 17, 2009   Volume 181, Issue 1 72-73 doi: 10.1016/j.tvjl.2009.03.014
McGreevy PD, McLean AN, Keay KA, Thomson PC.Trained responses are said to be under stimulus control when they appear reliably and exclusively on cue. The SMART system is a conceptual, three-dimensional graphic that uses four quadrants to chart a horse's responsiveness to various cues from two reins and the trainer's legs and seat. The current plots have been designed for ridden horses. The models assume that the trainer's cues are bilaterally equivalent, unless the intention is to signal a turn or a lateral movement. The extent to which 'go' signals manifest as straight, forward locomotion reflect the bilateral balance of the horse's re...
A preliminary study into rider asymmetry within equitation.
Veterinary journal (London, England : 1997)    April 16, 2009   Volume 181, Issue 1 34-37 doi: 10.1016/j.tvjl.2009.03.016
Symes D, Ellis R.Rider asymmetry is anecdotally associated with underperformance and injury. In this study, rider axial rotation (AR) and shoulder angle displacement (SAD) were measured using video analysis, and leg length inequality (LLI) was measured to assess any correlation between LLI, AR and SAD. Two video cameras, recording transverse and sagittal views, simultaneously filmed each of 17 female riders in walk, trot, and left and right canter. The rider's shoulder angles relative to the cranial caudal line were measured at each limb impact for a complete stride cycle. All riders exhibited left AR and a gr...
Use of kinetic gait analysis for detection, quantification, and differentiation of hind limb lameness and spinal ataxia in horses.
Journal of the American Veterinary Medical Association    March 3, 2009   Volume 234, Issue 5 644-651 doi: 10.2460/javma.234.5.644
Ishihara A, Reed SM, Rajala-Schultz PJ, Robertson JT, Bertone AL.To evaluate use of kinetic gait analysis for detection, quantification, and differentiation of hind limb lameness and spinal ataxia in horses. Methods: Prospective clinical study. Methods: 36 horses. Procedures-Kinetic gait analysis with a force plate was performed for 12 clinically normal horses, 12 horses with hind limb lameness, and 12 horses with spinal ataxia. Kinetic variables were compared among groups, correlated to subjective grading, and used to build predictive models to assess the accuracy of discrimination. Results: Subsets of kinetic variables were characteristically altered in a...
Use of a stand-alone pressure plate for the objective evaluation of forelimb symmetry in sound ponies at walk and trot.
Veterinary journal (London, England : 1997)    February 13, 2009   Volume 183, Issue 3 305-309 doi: 10.1016/j.tvjl.2008.12.012
Oosterlinck M, Pille F, Back W, Dewulf J, Gasthuys F.Subtle lameness in horses may be difficult to diagnose and methods to evaluate lameness objectively are useful when equine clinicians fail to reach a consensus. The aim of this study was to determine whether equine pressure plate measurements are repeatable when used to calculate forelimb loading (peak vertical pressure [PVP], peak vertical force [PVF], vertical impulse [VI]) and symmetry ratios, and to establish if these data are similar to the 'gold standard' force plate values. Since plate dimensions are relatively small, ponies were used to enable recordings to be taken from both forelimbs...
Accuracy of the TurfTrax Racing Data System for determination of equine speed and position.
Equine veterinary journal    January 24, 2009   Volume 40, Issue 7 680-683 doi: 10.2746/042516408x330338
Spence AJ, Tan H, Wilson A.The speed and position data collected by TurfTrax Racing Data Limited during UK Thoroughbred racing have potential to benefit equine science and welfare. The size (the 2006 data set alone consists of 30,932 individual horse starts across 2667 races) and nature (speed and 2D position for each horse at 4 updates per second) of the data make it a unique resource for questions in equine safety, welfare, performance, and animal locomotion. Objective: To determine the accuracy of the TurfTrax tracking system in estimating the speed and position of horses during racing. Methods: Measurements from the...
Design and validation of a dynamometric horseshoe for the measurement of three-dimensional ground reaction force on a moving horse.
Journal of biomechanics    January 10, 2009   Volume 42, Issue 3 336-340 doi: 10.1016/j.jbiomech.2008.11.017
Chateau H, Robin D, Simonelli T, Pacquet L, Pourcelot P, Falala S, Denoix JM, Crevier-Denoix N.Properties of ground surfaces condition locomotion, and quality of track surfaces is believed to be involved in the pathogenesis of many musculoskeletal injuries in the horse. Measuring ground reaction forces (GRF) is an interesting approach to assess those interactions. Forceplates are the most commonly used but they are not well suited to compare different ground surfaces at fast gaits. Embarked equipment, fixed under the horse's hoof, would allow force measurement on any track. The objective of this work was (1) to design a device which enables the measurement of 3-D GRF on any ground, at a...
Effects of trunk deformation on trunk center of mass mechanical energy estimates in the moving horse, Equus caballus.
Journal of biomechanics    December 23, 2008   Volume 42, Issue 3 308-311 doi: 10.1016/j.jbiomech.2008.11.005
Nauwelaerts S, Kaiser L, Malinowski R, Clayton HM.The estimation of the position of the center of mass (CM) is essential in a wide range of biomechanical analyses. In horses, the majority of the body mass is contained in the trunk and in most studies, the trunk is assumed to be rigid. However, this rigidity assumption has not been tested. We quantified changes in the position of the trunk CM due to external shape changes by measuring the kinematics of a mesh encompassing the trunk. Using a frame of reference fixed to the horse's spine, we described the shape deformation of the trunk during walking. In addition, we tested for speed and individ...
2D versus 3D in the kinematic analysis of the horse at the trot.
Veterinary research communications    December 11, 2008   Volume 33, Issue 6 507-513 doi: 10.1007/s11259-008-9196-x
Miró F, Santos R, Garrido-Castro JL, Galisteo AM, Medina-Carnicer R.The handled trot of three Lusitano Purebred stallions was analyzed by using 2D and 3D kinematical analysis methods. Using the same capture and analysis system, 2D and 3D data of some linear (stride length, maximal height of the hoof trajectories) and angular (angular range of motion, inclination of bone segments) variables were obtained. A paired Student T-test was performed in order to detect statistically significant differences between data resulting from the two methodologies With respect to the angular variables, there were significant differences in scapula inclination, shoulder angle, c...
Limits to running speed in dogs, horses and humans.
The Journal of experimental biology    December 2, 2008   Volume 211, Issue Pt 24 3836-3849 doi: 10.1242/jeb.024968
Denny MW.Are there absolute limits to the speed at which animals can run? If so, how close are present-day individuals to these limits? I approach these questions by using three statistical models and data from competitive races to estimate maximum running speeds for greyhounds, thoroughbred horses and elite human athletes. In each case, an absolute speed limit is definable, and the current record approaches that predicted maximum. While all such extrapolations must be used cautiously, these data suggest that there are limits to the ability of either natural or artificial selection to produce ever fast...
Stabilization and mobility of the head, neck and trunk in horses during overground locomotion: comparisons with humans and other primates.
The Journal of experimental biology    December 2, 2008   Volume 211, Issue Pt 24 3889-3907 doi: 10.1242/jeb.020578
Dunbar DC, Macpherson JM, Simmons RW, Zarcades A.Segmental kinematics were investigated in horses during overground locomotion and compared with published reports on humans and other primates to determine the impact of a large neck on rotational mobility (> 20 deg.) and stability (< or = 20 deg.) of the head and trunk. Three adult horses (Equus caballus) performing walks, trots and canters were videotaped in lateral view. Data analysis included locomotor velocity, segmental positions, pitch and linear displacements and velocities, and head displacement frequencies. Equine, human and monkey skulls and cervical spines were measured to es...
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