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Topic:Cortisol

Cortisol is a steroid hormone produced by the adrenal glands in horses, playing a significant role in the regulation of metabolism, immune response, and stress management. As a glucocorticoid, cortisol influences various physiological processes, including glucose metabolism, inflammation modulation, and the maintenance of homeostasis during stress. Its levels in horses can vary due to factors such as exercise, environmental changes, and psychological stressors. Cortisol is often measured to assess stress responses and overall well-being in equine health management. This page compiles peer-reviewed research studies and scholarly articles that explore the production, regulation, and implications of cortisol in equine physiology and health.
Immunological response to long-term transport stress in mature horses and effects of adaptogenic dietary supplementation as an immunomodulator.
Equine veterinary journal    December 8, 2004   Volume 36, Issue 7 583-589 doi: 10.2746/0425164044864589
Stull CL, Spier SJ, Aldridge BM, Blanchard M, Stott JL.Little information exists on the immunological effects of transport or the use of supplements to minimise transport stress. Objective: To establish baseline ranges and evaluate immunophenotypic and functional changes associated with transport and a nutritional 'adaptogen' supplement. Methods: Horses received either supplement (n = 10) or placebos (n = 9) during the 30 day study. After 28 days in stalls, 12 horses (6 supplement; 6 placebo) were transported for 24 h, then unloaded and recovered. Venous blood samples were collected on Days 1, 14 and 28 to establish baselines, and on Days 28, 29 a...
Effect of chronic clenbuterol administration and exercise training on immune function in horses.
Journal of animal science    November 13, 2004   Volume 82, Issue 12 3500-3507 doi: 10.2527/2004.82123500x
Malinowski K, Kearns CF, Guirnalda PD, Roegner V, McKeever KH.Effects of longitudinal exercise training and acute intensive exercise (simulated race test) on immune function have not been reported in horses. Clenbuterol, a beta2-adrenergic agonist, is used to manage inflammatory airway disease in horses. This study investigated the interaction of 8 wk of exercise training with or without 12 wk of clenbuterol administration in horses. Twenty-three untrained standardbred mares (10 +/- 3 yr, Mean +/- SE) were used and divided into four experimental groups. Horses given clenbuterol plus exercise (CLENEX; n = 6) and clenbuterol alone (CLEN; n = 6) received 2....
Biological availability of inhaled fluticasone propionate in horses.
The Veterinary record    October 21, 2004   Volume 155, Issue 12 361-364 doi: 10.1136/vr.155.12.361
Laan TT, Westermann CM, Dijkstra AV, van Nieuwstadt RA, Fink-Gremmels J.Healthy horses received aerosolised, intranasal or oral doses of 3 mg of fluticasone propionate evenly divided over morning and evening treatments for seven days. The bioavailability of the drug was determined in terms of the suppression of the endogenous cortisol concentrations in the horses during the period of treatment. The horses which received the aerosolised drug had significantly lower concentrations of endogenous cortisol on days 5 and 8 than the horses which received aerosolised placebo. The horses which received nasal and oral doses of fluticasone propionate showed no significant ch...
Effects of continuous rate intravenous infusion of butorphanol on physiologic and outcome variables in horses after celiotomy.
Journal of veterinary internal medicine    August 24, 2004   Volume 18, Issue 4 555-563 doi: 10.1892/0891-6640(2004)182.0.co;2
Sellon DC, Roberts MC, Blikslager AT, Ulibarri C, Papich MG.A randomized, controlled, blinded clinical trial was performed to determine whether butorphanol administered by continuous rate infusion (CRI) for 24 hours after abdominal surgery would decrease pain and surgical stress responses and improve recovery in horses. Thirty-one horses undergoing exploratory celiotomy for abdominal pain were randomly assigned to receive butorphanol CRI (13 microg/kg/h for 24 hours after surgery; treatment) or isotonic saline (control). All horses received flunixin meglumine (1.1 mg/kg IV q12h). There were no significant differences between treatment and control horse...
The association between heart rate, heart rate variability, endocrine and behavioural pain measures in horses suffering from laminitis.
Journal of veterinary medicine. A, Physiology, pathology, clinical medicine    August 19, 2004   Volume 51, Issue 5 218-225 doi: 10.1111/j.1439-0442.2004.00627.x
Rietmann TR, Staᆲher M, Bernasconi P, Auer JA, Weishaupt MA.The objective of this study was to compare the stress response of horses suffering from laminitis after short- and long-term treatment with the intent to evaluate power spectral analysis of heart rate variability (HRV) for pain monitoring. Data were collected from 19 horses with acute or chronic exacerbating laminitis without known primary disease before and after treatment with non-steroidal anti-inflammatory drugs (NSAID). Recordings were carried out the day after admission to the equine hospital. Measurements were repeated on day 7 of the treatment. The recorded parameters included a clinic...
Serum insulin concentrations in horses with equine Cushing’s syndrome: response to a cortisol inhibitor and prognostic value.
Equine veterinary journal    May 19, 2004   Volume 36, Issue 3 295-298 doi: 10.2746/0425164044877288
McGowan CM, Frost R, Pfeiffer DU, Neiger R.Serum insulin concentration and its use as a prognostic indicator in horses with equine Cushing's syndrome (ECS) have been poorly documented. Objective: To examine daily insulin variations in horses with ECS and the effect of treatment using trilostane, a competitive inhibitor of 3beta-hydroxysteroid dehydrogenase. Further, we aimed to examine the relationship between baseline serum insulin concentration and survival in horses with ECS. Methods: Basal serum insulin concentrations were measured in 20 confirmed ECS cases by taking blood at regular 4 h intervals for 24 h (1200, 1600, 2000, 2400, ...
Tissue-specific dysregulation of cortisol metabolism in equine laminitis.
Equine veterinary journal    February 6, 2004   Volume 36, Issue 1 41-45 doi: 10.2746/0425164044864750
Johnson PJ, Ganjam VK, Slight SH, Kreeger JM, Messer NT.The role of glucocorticoids (GCs) in the pathogenesis of laminitis is incompletely understood. Local tissue activity of GC is regulated by the steroid converting enzyme, 11beta-hydroxysteroid dehydrogenase-1 (11beta-HSD-1). Changes in integumentary (skin and hoof lamellar) 11beta-HSD activity occurring during laminitis could affect the extent to which GCs are involved in its development. Objective: That changes in integumentary 11beta-HSD-1 activity associated with the laminitic condition would lead to elevated local tissue levels of GCs, which could subsequently contribute, through paracrine ...
Effects of midazolam on equine innate immune response: a flow cytometric study.
Veterinary immunology and immunopathology    September 13, 2003   Volume 95, Issue 1-2 11-19 doi: 10.1016/s0165-2427(03)00097-7
Massoco C, Palermo-Neto J.Benzodiazepines (BDZ) are among the most frequently used class of psychotropic drugs employed in veterinary medicine in Brazil and worldwide due to their anxiolytic, muscle relaxant and anticonvulsant effects [J. Clin. Pharmacol. 33 (1993) 124]. Peripheral benzodiazepine receptor (PBR) sites were described in peripheral organs, endocrine steroidogenic tissues and immune organs and cells. Midazolam is a mixed-type agonist of PBRs. The present study is focused on the effects of midazolam on equine peripheral blood neutrophils, peritoneal macrophages and cortisol levels in plasma. Adult horses we...
Endocrine responses in mares and geldings with high body condition scores grouped by high vs. low resting leptin concentrations.
Journal of animal science    September 13, 2003   Volume 81, Issue 9 2311-2321 doi: 10.2527/2003.8192311x
Cartmill JA, Thompson DL, Storer WA, Gentry LR, Huff NK.Previous observations from this laboratory indicated that horses with high BCS could have resting plasma leptin concentrations ranging from low (1 to 5 ng/mL) to very high (10 to 50 ng/mL). To study the possible interactions of leptin secretion with other endocrine systems, BCS and plasma leptin concentrations were measured on 36 mares and 18 geldings. From mares and geldings that had a mean BCS of at least 7.5, five with the lowest (low leptin) and five with the highest (high leptin) leptin concentrations were selected. Jugular blood samples were collected twice daily for 3 d from the 20 sele...
Comparison of efficacy and tolerability of isoflupredone and dexamethasone in the treatment of horses affected with recurrent airway obstruction (‘heaves’).
Equine veterinary journal    July 26, 2003   Volume 35, Issue 4 419-424 doi: 10.2746/042516403776014208
Picandet V, Léguillette R, Lavoie JP.Corticosteroids are currently the most effective drugs for the control of 'heaves' in horses. However, there is limited information concerning the comparative efficacy and tolerability of the various corticosteroids when used for treatment. Objective: To compare the therapeutic and side effects of isoflupredone acetate to those of dexamethasone. Methods: A parallel design to compare the effects of 2 corticosteroids by evaluating lung function, serum cortisol and electrolyte concentrations, response to ACTH stimulation and haematology sequentially during a 14 day control period (no treatment), ...
Efficacy of trilostane for the treatment of equine Cushing’s syndrome.
Equine veterinary journal    July 26, 2003   Volume 35, Issue 4 414-418 doi: 10.2746/042516403776014271
McGowan CM, Neiger R.Trilostane, a competitive 3-beta hydroxysteroid dehydrogenase inhibitor, has been used successfully to control clinical signs and cortisol excess in canine pituitary dependent hyperadrenocorticism. Objective: Trilostane was evaluated for its efficacy in resolving clinical and clinicopathological abnormalities of equine Cushing's syndrome (ECS) and to assess its safety. Methods: Twenty horses (mean age 21 years) diagnosed with ECS were followed for 1 or 2 years. Affected horses received 0.4-1 mg/kg (mean 0.5 mg/kg) trilostane once daily. Results: Clinical signs assessed over 1 or 2 years, showe...
Glucose clearance in grazing mares is affected by diet, pregnancy, and lactation.
Journal of animal science    July 12, 2003   Volume 81, Issue 7 1764-1771 doi: 10.2527/2003.8171764x
Hoffman RM, Kronfeld DS, Cooper WL, Harris PA.The glucose tolerance test in the horse may be used to determine metabolic responses to diet, disease, or physiologic state. The objective of this study was to determine the effect of reproductive stage (gestation and lactation) and supplemental dietary energy source (sugar and starch [SS] or fiber and fat [FF]) on glucose metabolism in grazing mares using an oral glucose tolerance test. Twelve mares, six on each supplement, were examined on three occasions: one in the third trimester of pregnancy, the second in early lactation, and the third in late lactation. During each test, venous samples...
Prolonged suppression of the innate immune system in the horse following an 80 km endurance race.
Equine veterinary journal    March 18, 2003   Volume 35, Issue 2 133-137 doi: 10.2746/042516403776114144
Robson PJ, Alston TD, Myburgh KH.An increased susceptibility to bacterial and viral infections of the respiratory tract, which results in a loss of performance, has been reported in racehorses. Much research has focused on the influence of high-intensity exercise of a short duration on immune system function in horses, but scant attention has been given to prolonged endurance exercise as an immune modulator. Objective: The objective of this study was to evaluate the effect of an 80 km endurance race on the monocyte and neutrophil oxidative burst, serum cortisol, glutamine and plasma glucose concentrations in 8 endurance-train...
Developmental changes in pulmonary and renal angiotensin-converting enzyme concentration in fetal and neonatal horses.
Reproduction, fertility, and development    March 6, 2003   Volume 14, Issue 7-8 413-417 doi: 10.1071/rd02044
O'Connor SJ, Fowden AL, Holdstock N, Giussani DA, Forhead AJ.Angiotensin-converting enzyme (ACE) has an active role in the control of blood pressure and body fluid homeostasis both before and after birth. This study investigated the ontogeny of pulmonary and renal ACE concentrations in fetal and neonatal horses. Fetal pulmonary ACE concentration increased from 250 days towards term (c. 335 days). Newborn foals showed significantly higher mean concentrations of pulmonary ACE (4.40 +/- 0.62 nmol min(-1) mg protein(-1)) than both fetuses during late gestation (1.23 +/- 0.51 nmol min(-1) mg protein(-1)) and animals aged 1 day to 2 weeks of postnatal age (0....
Plasma adrenocorticotropin (ACTH) concentrations and clinical response in horses treated for equine Cushing’s disease with cyproheptadine or pergolide.
Equine veterinary journal    November 29, 2002   Volume 34, Issue 7 679-685 doi: 10.2746/042516402776250333
Perkins GA, Lamb S, Erb HN, Schanbacher B, Nydam DV, Divers TJ.Plasma ACTH levels have been variable in horses with a positive clinical response for therapy for equine Cushing's Disease (ECD). Therefore, our purpose was to determine the value of monitoring plasma adrenocorticotropin (ACTH) levels during treatment of equine Cushing's disease (ECD) with either cyproheptadine (n = 32) or pergolide (n = 10). First, we validated the chemiluminescent ACTH assay (specificity, precision, accuracy, intra-assay and interassay variations) and tested methods of handling the whole blood from the time of collection to when the ACTH was assayed. The sensitivity and spec...
Effects of dexamethasone, glucose infusion, adrenocorticotropin, and propylthiouracil on plasma leptin concentrations in horses.
Domestic animal endocrinology    November 27, 2002   Volume 24, Issue 1 1-14 doi: 10.1016/s0739-7240(02)00183-2
Cartmill JA, Thompson DL, Gentry LR, Pruett HE, Johnson CA.In experiment 1, nine light horse geldings (three 3 x 3 Latin squares) received dexamethasone (DEX; 125 microg/kg BW, i.m.), glucose (0.2 g/kg BW, i.v.), or nothing (control) once per day for 4 days. DEX increased (P < 0.001) glucose, insulin, and leptin concentrations and resulted in a delayed increase (P < 0.001) in IGF-I concentrations. In experiment 2, mares were similarly treated with DEX (n = 6) or vehicle (n = 6). DEX again increased (P < 0.01) glucose, insulin, and leptin concentrations; the delayed elevation in IGF-I concentrations occurred on day 10, 12, and 19, relative to ...
Postnatal cardiovascular function after manipulation of fetal growth by embryo transfer in the horse.
The Journal of physiology    November 15, 2002   Volume 547, Issue Pt 1 67-76 doi: 10.1113/jphysiol.2002.027409
Giussani DA, Forhead AJ, Gardner DS, Fletcher AJ, Allen WR, Fowden AL.This study used between-breed embryo transfer in the horse to investigate the effects of maternal size and uterine capacity on fetal growth and postnatal cardiovascular and neuroendocrine functions. Equine embryos were transferred to establish eight Thoroughbred-in-Thoroughbred (TinT), seven Pony-in-Pony (PinP), five Thoroughbred-in-Pony (TinP) and eight Pony-in-Thoroughbred (PinT), pregnancies. Maternal and foal weights and placental microscopic area were measured at birth. At 6 days of postnatal life, arterial blood pressure and heart rate were monitored and blood samples were taken for horm...
Speed associated with plasma pH, oxygen content, total protein and urea in an 80 km race.
Equine veterinary journal. Supplement    October 31, 2002   Issue 34 39-43 doi: 10.1111/j.2042-3306.2002.tb05389.x
Hoffman RM, Hess TM, Williams CA, Kronfeld DS, Griewe-Crandell KM, Waldron JE, Graham-Thiers PM, Gay LS, Splan RK, Saker KE, Harris PA.To test the hypothesis that endurance performance may be related quantitatively to changes in blood, we measured selected blood variables then determined their reference ranges and associations with speed during an 80 km race. The plan had 46 horses in a 2 x 2 factorial design testing a potassium-free electrolyte mix and a vitamin supplement. Blood samples were collected before the race, at 21, 37, 56 and 80 km, and 20 min after finishing, for assay of haematocrit, plasma pH, pO2, pCO2, [Na+], [K+], [Ca++], [Mg++], [Cl-], lactate, glucose, urea, cortisol, alpha-tocopherol, ascorbate, creatine ...
Metabolic and osmoregulatory function at low and high (3800 m) altitude.
Equine veterinary journal. Supplement    October 31, 2002   Issue 34 545-550 doi: 10.1111/j.2042-3306.2002.tb05480.x
Greene HM, Cogger EA, Miltenberger TL, Koch AK, Bray RE, Wickler SJ.Altitude evokes physiological adjustments that include not only respiratory and cardiovascular properties, but also metabolic function, renal and endocrine responses. The purpose of the present study was designed to expand our understanding of the physiological process involved with acclimatisation to high altitude in equids. The study examined temporal effects on metabolic and osmoregulatory function in horses (n = 6) at rest and postexercise at 3800 m. Animals were studied at 225 m (Pb = 743 mmHg) and during a 10 day stay at altitude (Pb = 487 mmHg). Rest samples were taken 90 min postprandi...
Effects of cool and hot humid environmental conditions on neuroendocrine responses of horses to treadmill exercise.
Veterinary journal (London, England : 1997)    October 3, 2002   Volume 164, Issue 1 54-63 doi: 10.1053/tvjl.2002.0721
Williams RJ, Marlin DJ, Smith N, Harris RC, Haresign W, Davies Morel MC.To determine the effects of exercise, high heat and humidity and acclimation on plasma adrenaline, noradrenaline, beta-endorphin and cortisol concentrations, five horses performed a competition exercise test (CET; designed to simulate the speed and endurance test of a three-day event) in cool dry (CD) (20 degrees C/40% RH) and hot humid (30 degrees C/80% RH) conditions before (pre-acclimation) and after (post-acclimation) a 15 day period of humid heat acclimation. Plasma adrenaline and noradrenaline concentrations pre-acclimation were significantly increased compared with exercise in the CD tr...
Effects of cross-tying horses during 24 h of road transport.
Equine veterinary journal    October 3, 2002   Volume 34, Issue 6 550-555 doi: 10.2746/042516402776180214
Stull CL, Rodiek AV.Transportation stress has been implicated as a predisposing factor to respiratory disease in horses. Cross-tying horses individually in stalls is common practice for transporting show and racehorses, but horses also travel in small groups or individually without being restricted by tying. The objective of this study was to compare physiological responses of horses travelling cross-tied or loose during 24 h of road transport. Ten horses were used in a cross-over design consisting of two 4 day trials. In the first trial, 6 horses were cross-tied, while 2 pairs of horses were loose in enclosed co...
Urinary cortisol:creatinine ratios in healthy horses and horses with hyperadrenocorticism and non-adrenal disease.
The Veterinary record    July 24, 2002   Volume 150, Issue 25 773-776 doi: 10.1136/vr.150.25.773
Chandler KJ, Dixon RM.Urinary cortisol and creatinine concentrations, and the cortisol:creatinine ratio were compared between 12 healthy horses (group 1), 13 horses with Cushing's disease (group 2), and eight horses with dysautonomia syndrome (equine grass sickness) (group 3). The mean (sd) urinary cortisol concentrations were 112 (55.7), 250 (357) and 864 (526) nmol/litre in groups 1, 2 and 3, respectively; the mean (sd) urinary creatinine concentrations were 18.9 (7.3), 12.0 (6.7) and 45.2 (26.4) nmol/litre in groups 1, 2 and 3, respectively, and the mean (sd) ratios were 6.1 (2.6), 19.8 (23.8) and 21.3 (14.5) (x...
Exercise-induced changes in atrial peptides in relation to neuroendocrine responses and fluid balance in the horse.
Journal of veterinary medicine. A, Physiology, pathology, clinical medicine    May 22, 2002   Volume 49, Issue 3 144-150 doi: 10.1046/j.1439-0442.2002.00428.x
Kokkonen UM, Pösö AR, Hyyppä S, Huttunen P, Leppäluoto J.Previous data show that, in horses, plasma atrial natriuretic peptides (ANP and NT-ANP) remain elevated for a long time after exercise. To study whether exercise-induced changes in hormonal and fluid balance explain this, we measured plasma concentrations of COOH- and NH2-terminal atrial natriuretic peptides (ANP(99-129) and NT-ANP(1-98) together with arginine vasopressin (AVP), adrenocorticotrophin (ACTH), beta-endorphin, cortisol, catecholamines, and indicators of fluid balance in six Finnhorses after a graded submaximal exercise test on a treadmill. After exercise, AVP and catecholamines di...
Effects of racing on lymphocyte proliferation in horses.
American journal of veterinary research    April 10, 2002   Volume 63, Issue 4 528-530 doi: 10.2460/ajvr.2002.63.528
Nesse LL, Johansen GI, Blom AK.To measure the lymphocyte proliferation response in horses 12 to 16 hours after completion of a race. Methods: 8 Thoroughbreds that competed in 14 races and 3 control Thoroughbreds that did not race. Methods: Horses participated in races during the late afternoon or evening. Venous blood samples were collected on a morning before a race (1 or 2 days before the race or on the day of the race), on the afternoon of a race (40 to 60 minutes after the race), and on the morning of the day after a race (12 to 16 hours after the race). Lymphocyte proliferation responses and WBC count were measured in ...
Effectiveness of a two-dose regimen of prostaglandin administration in inducing luteolysis without adverse side effects in mares.
Equine veterinary journal    March 22, 2002   Volume 34, Issue 2 191-194 doi: 10.2746/042516402776767240
Irvine CH, McKeough VL, Turner JE, Alexander SL, Taylor TB.Our objectives were to determine whether repeated administration of prostaglandin F2alpha (PGF2alpha) to simulate the endogenous mode of secretion would be more effective than a single injection in inducing luteolysis and enable use of smaller doses less likely to cause adverse side effects. The main study comprised 43 dioestrous mares, who were given im. either a single 10 mg dose of natural PGF2alpha (n = 22) or 2 doses of 0.5 mg PGF2, 24 h apart (n = 21). The intensity of side effects was assessed in 8 dioestrous mares given 5, 1.5, 0.5 or 0 mg PGF2alpha in consecutive cycles. Two doses of ...
Growth hormone response to a novel growth hormone-releasing tripeptide in horses: interaction with gonadotropin-releasing hormone, thyrotropin-releasing hormone, and sulpiride.
Journal of animal science    March 14, 2002   Volume 80, Issue 3 744-750 doi: 10.2527/2002.803744x
Kennedy SR, Thompson DL, Pruett HE, Burns PJ, Deghenghi R.A series of experiments was performed to determine the factor(s) responsible for an apparent inhibition of GH secretion in mares administered the GH secretagogue EP51389 in combination with GnRH, thyrotropin-releasing hormone (TRH), and sulpiride. Experiment 1 tested the repeatability of the original observation: 10 mares received EP51389 at 10 microg/kg BW; five received TRH (10 microg/kg BW), GnRH (1 microg/kg BW), and sulpiride (100 microg/kg BW) immediately before EP51389, and five received saline. The mixture of TRH, GnRH, and sulpiride reduced (P = 0.0034) the GH response to EP51389, con...
The role of nutritional therapy in the treatment of equine Cushing’s syndrome and laminitis.
Alternative medicine review : a journal of clinical therapeutic    October 10, 2001   Volume 6 Suppl S4-S16 
Harman J, Ward M.Equine Cushing's syndrome, a relatively common and complex condition, is difficult to treat with conventional medicine. Cushing's syndrome involves a hyperplasia or adenoma of the anterior pituitary gland. Biochemical alterations include increased endogenous cortisol, insulin resistance, elevated adrenocorticotrophic hormone, and decreased thyroid hormone levels. Symptoms include hirsutism with no loss of the winter coat in summer, refractory laminitis, weight problems (over- or underweight), polyuria/polydipsia (Pu/Pd), frequent infections, lowered immunity to intestinal parasites, decreased ...
Effects of nandrolone treatment on recovery in horses after strenuous physical exercise.
Journal of veterinary medicine. A, Physiology, pathology, clinical medicine    September 14, 2001   Volume 48, Issue 6 343-352 doi: 10.1046/j.1439-0442.2001.00368.x
Hyyppä S.To test the effect of nandrolone on their recovery, six adult half-bred riding horses performed a competition exercise test (CET) and a standardized exercise test (SET) on consecutive days before and after a 2-week treatment with the anabolic steroid nandrolone laurate. Blood samples were collected during and between these tests for the determination of red cell volume and concentrations of blood lactate, plasma glucose, non-esterified fatty acids, glycerol, triglycrides, erythropoietin, cortisol, insulin, and glucagon. Muscle biopsy specimens were taken immediately after the CET and before th...
Salivary and plasma concentration of cortisol in normal horses and horses with Cushing’s disease.
Equine veterinary journal    March 27, 2001   Volume 33, Issue 2 211-213 doi: 10.1111/j.2042-3306.2001.tb00604.x
van der Kolk JH, Nachreiner RF, Schott HC, Refsal KR, Zanella AJ.No abstract available
Production of reactive oxygen species in neutrophils after repeated bouts of exercise in standardbred trotters.
Journal of veterinary medicine. A, Physiology, pathology, clinical medicine    March 14, 2001   Volume 47, Issue 9 565-573 doi: 10.1046/j.1439-0442.2000.00275.x
Korhonen PA, Lilius EM, Hyyppä S, Räsänen LA, Pösö AR.Six trained Standardbred trotters exercised on a racetrack on 2 days with a 3-day interval. On both exercise days the horses trotted three different exercise bouts with increasing intensity with 60-min intervals. Exercise-induced stress was manifested as leucocytosis, an increase in the neutrophil:lymphocyte (N:L) ratio, and increased capacity to produce reactive oxygen species in the peripheral blood as indicated by an increase in whole blood chemiluminescence. The leucocytosis was mainly due to neutrophilia, which lasted for 6 h. Production of reactive oxygen species per single neutrophil sh...
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