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Topic:Equids

Equids are members of the family Equidae, which includes modern horses, donkeys, and zebras, as well as extinct species. Equids are characterized by their long limbs, single-toed hooves, and herbivorous diet, which is primarily composed of grasses. Horses, specifically, have been domesticated for thousands of years and have played significant roles in agriculture, transportation, and sport. This page compiles peer-reviewed research studies and scholarly articles that explore various aspects of equids and horses, including their evolution, physiology, behavior, and interactions with humans. The collected works provide insights into the genetic diversity, adaptive traits, and conservation efforts related to equid species.
Equine cyathostomins-Hidden histories.
Advances in parasitology    June 26, 2026   Volume 133 1-25 doi: 10.1016/bs.apar.2026.06.002
Mulcahy G, Byrne O, Walshe N.Cyathostomins (also known as small strongyles or small redworms) are ubiquitous parasites of equids. They are found in a wide range of climatic regions, and inhabit the large intestine of their hosts. While infections are often sub-clinical, clinical consequences ranging from chronic to acute, and most notably, acute larval cyathostominosis, can occur in specific circumstances. Control of cyathostomin infection is complicated by widespread drug resistance. This Chapter provides a broad review of cyathostomin biology and host-parasite relationships, and also highlights significant knowledge gap...
Computed tomography long-term follow-up of a facial fracture involving the orbit and paranasal sinus.
Equine veterinary journal    June 24, 2026   doi: 10.1002/evj.70224
Kalinovskiy A, Lischer C, Eule JC, Thieme K, Mählmann K.Fractures of the facial bones involving the orbit are challenging conditions that require surgical treatment. Objective: To describe preoperative and postoperative clinical and computed tomographic findings of an orbital rim fracture repaired using a reconstruction plate. Methods: Case report. Methods: A 14-year-old Haflinger mare was presented with an open, depressed and comminuted fracture involving the frontal, nasal, lacrimal and maxillary bone, sinus system and the medial aspect of the orbital rim and wall on the right side, as diagnosed by CT. Fracture repair included orbital rim reconst...
Comments on ‘Finite element modelling for investigating bone-implant interactions in equine orthopaedics’.
Equine veterinary journal    June 21, 2026   doi: 10.1002/evj.70221
de Souza AF, Meira JBC, Fonteque JH, De Zoppa ALDV.No abstract available
Circulating Elements in the Bloodstream of Growing Foals: Focus on Heavy Metals and Bone Remodeling Across Growth Stages.
Biological trace element research    June 19, 2026   doi: 10.1007/s12011-026-05186-3
Hulitká D, Coradduzza D, Sanna A, Biggio GP, Taras A, Cannas A, Carru C, Medici S, Cherchi R, Cappai MG.Bone tissue undergoes continuous remodelling throughout the life of the individual, and growth is the most impacting phase. In fact, tightly coordinated processes of resorption, synthesis, and mineralization of the bone matrix under systemic hormonal, biomechanical, and metabolic control occur for variable periods of time across species. The horse is a fast-growing animal and for this reason identified as precocial. It was hypothesized that blood serum levels of minerals in growing foals at different stages of accretion may vary over time. This investigation aimed to assess the dynamics of cir...
Matched embryo-endometrium RNA-seq reveals coordinated but asymmetric transcriptomic reprogramming at the onset of early equine pregnancy.
Frontiers in cell and developmental biology    June 18, 2026   Volume 14 1840498 doi: 10.3389/fcell.2026.1840498
Da Silva Álvarez E, Ortiz-Rodríguez JM, Martín-Cano FE, Álvarez-Barrientos A, Becerro-Rey L, Gil MC, Redondo E, Masot J, Aparicio IM, Peña FJ....Maternal recognition of pregnancy (MRP) in the mare requires coordinated adaptations of both the conceptus and the endometrium during the peri-recognition interval; however, integrated transcriptomic analyses of both compartments within the same biological window remain limited. Here, we analyzed conceptus RNA-seq data across embryonic age (Days 8 and 12 post ovulation) and endometrial RNA-seq data across pregnancy status (pregnant versus non-pregnant mares sampled between Days 8 and 12 post ovulation). In pregnant mares, conceptus recovery and endometrial biopsy collection were performed with...
Differences in Stride Characteristics Between Lead and Wheel Horses in Competitive Chuckwagon Racing.
Animals : an open access journal from MDPI    June 18, 2026   Volume 16, Issue 12 1890 doi: 10.3390/ani16121890
Pfau T, Broek MVD, Davis BL, De Bruyne C, Eamon CM, Fallscheer M, Frostad S, Garcia-Alamo K, Skotarek Loch S, Takahashi Y, Weller R, Chan ZYS.In Chuckwagon racing, teams of four Thoroughbreds with two lead horses in front and two wheel horses directly behind pull a 600 kg wagon around a racetrack. Our study focuses on stride parameters that have previously retrospectively predicted impending injuries. We hypothesize that in aid of the initiation of the tight figure-of-eight turn at the start of the race, the more agile lead horses gallop with higher stride frequency (SF) and reduced stride length (SL), and that differences are exacerbated in the initial accelerative race segments. Methods: Speed, SF, and SL were quantified with glob...
Transcriptomic and Meat Quality Differences in Longissimus Dorsi Muscle of Surgically Castrated Three-Year-Old Kazakh Horses.
Biology    June 18, 2026   Volume 15, Issue 12 959 doi: 10.3390/biology15120959
Li Z, Ren W, Wang R, Li L, Ma S, Su Y, Shan D, Huang Q, Wang J.Although the Kazakh horse is a dual-purpose breed renowned for both milk and meat production, the extent to which surgical castration alters gene expression in its muscles has not yet been fully elucidated. In this study, left longissimus dorsi muscle (LDM) samples were obtained from six Kazakh stallions (W group) and six Kazakh geldings (S group) to comparatively evaluate meat quality parameters, examine histological characteristics in tissue sections, and apply transcriptomic profiling to comprehensively explore the principal regulatory pathways and candidate genes through which surgical cas...
A Comprehensive Analysis of the Agreement and Performance of Variant Annotation Programs in Equine Genomes.
Genes    June 18, 2026   Volume 17, Issue 6 704 doi: 10.3390/genes17060704
Marlowe JL, Hughes L, Barrey E, Leeb T, Bellone R, McCue ME, Durward-Akhurst S. Advances in whole-genome sequencing (WGS) technology have led to the widespread adoption of WGS for investigating genetic diseases and traits in domestic animals. This has created a need for improved methods for prioritizing candidate causal variants. One way variants are prioritized is using variant annotators that predict variant effects based on their proximity to genomic features and effect on amino acid sequence. However, validation of variant annotators for domestic animal genomes is lacking. In this study, we calculated the agreement of three popular variant annotators, Ensembl Varian...
Redefining stallion sperm bioenergetics: lessons from a decade of change in mammalian sperm metabolism.
Reproduction (Cambridge, England)    June 17, 2026   Volume 172, Issue 1 xaag077 doi: 10.1093/reprod/xaag077
Peña FJ, Becerro-Rey L, Ortega-Ferrusola C, da Silva-Álvarez E, Zabalo-Palomo C, Gil MC, Martín-Cano FE.In brief: Sperm bioenergetics is now understood as an integrated system in which ATP production, redox homeostasis, substrate availability, and cellular heterogeneity jointly determine sperm function. This review uses the stallion as an oxidative-biased model to synthesize advances from 2016 to 2026 and proposes a metabolically plastic framework with direct implications for semen preservation and fertility assessment. Abstract: Over the last decade, mammalian sperm metabolism has moved from a binary view centered on glycolysis versus oxidative phosphorylation to an integrated model in which AT...
Phenotypic characterization through identification of gait and coat color phenotype of the Mountain Pleasure Horse.
Journal of equine veterinary science    June 17, 2026   106066 doi: 10.1016/j.jevs.2026.106066
Irons A, Culwell J, Nicodemus MC, Vandiver M, North E, Cavinder C, Rios de Alvarez L, Williams T.Phenotypic characterization of traits like gait and coat color is useful for minor breed identification and conservation. The Mountain Pleasure Horse (MPH) is a minor American breed as it is listed as threatened by the Livestock Conservancy. Objective: The study objective was to utilize phenotypic characterization for identifying gait and coat color phenotype of the MPH. Methods: Gait was characterized utilizing frame-by-frame analysis to identify spatiotemporal variables of purebred horses (n = 6) performing the flat walk and saddle rack. Coat color was characterized using coat color assi...
Mastication characterization of horses fed bermudagrass hay in a two-part field study.
Journal of equine veterinary science    June 17, 2026   106068 doi: 10.1016/j.jevs.2026.106068
Long JF, Hutton R, Abrams A, Duberstein JA, Duberstein KJ.Bermudagrass (Cynodon dactylon L.) is a common forage in the southeastern United States but remains controversial in equine diets due to reported associations with impaction colic. Reduced fiber digestibility has been proposed as a contributing factor, however differences in mastication of fine-stemmed bermudagrass hay have not been studied and may play a role in digestive disturbances. Objective: To compare chewing characteristics, including chews per unit weight and chews per minute, in horses consuming bermudagrass hay compared to other common hay types. Methods: Two studies were conducted....
Glandular gastric disease prevalence in equids (horses, donkeys, and mules) and other gastric findings: post-mortem comparative study.
Veterinary research forum : an international quarterly journal    June 15, 2026   Volume 17, Issue 6 373-380 doi: 10.30466/vrf.2025.2056590.4709
Medina Bolivar AL, Faleiros RR, Martinez Aranzales JR.Equine glandular gastric disease (EGGD) has limited prevalence studies and scarce understanding of the predisposing factors and pathophysiological mechanisms involved. The objectives of this study were to determine and compare the post-mortem prevalence of EGGD and other gastric lesions in Colombian equids, specifically horses, donkeys, and mules. The study utilized a sample of 300 equids stomachs, with 100 specimens from each group, collected immediately after slaughter. The post-mortem examination included a thorough inspection of the ventral fundus, adjacent area of the margo plicatus, and ...
A Case of Separate os tarsale primum in a Horse.
Veterinary sciences    June 15, 2026   Volume 13, Issue 6 582 doi: 10.3390/vetsci13060582
Vladova D, Kostov D, Hristov H, Goranov N, Dinev T, Lioteva-Dineva A, Stamatova-Yovcheva K.Anatomical variation in the equine tarsus may involve either fusion of the with the or the presence of a separate . The distal row of tarsal bones is therefore of particular anatomical interest, as an independent first tarsal bone has been only sporadically documented in previous studies. In this study, twelve equine tarsal joints were examined using macroscopic and radiographic methods. In eleven specimens, six tarsal bones were identified, whereas in one specimen (1/12, 8.3%), seven tarsal bones were observed due to the presence of an independent with an irregular cuboid shape. Radiograph...
Prevalence and Genetic Characterization of Giardia duodenalis and Blastocystis sp. in Horses in Shanxi Province, North China.
Animals : an open access journal from MDPI    June 15, 2026   Volume 16, Issue 12 1845 doi: 10.3390/ani16121845
Liu XZ, Su N, Li W, Wang DY, Zhang ZD, Zhu XQ, Gao WW. (synonyms and ) and sp. are prevalent zoonotic intestinal protozoans with significant public health and economic importance. Both parasites infect various hosts, including humans and most livestock. can cause infections ranging from asymptomatic carriage to clinical manifestations such as acute diarrheal disease, growth stunting and malabsorption, whereas sp. infection is often asymptomatic and its pathogenicity remains debated. Following the rapid expansion of modern equine industries (horse racing, equestrian sports, production, and tourism), concern about the spread of parasitic diseas...
Prevalence and spatial distribution of Brucella spp. seropositive equids from the state of Mato Grosso, Brazil.
Journal of equine veterinary science    June 13, 2026   106064 doi: 10.1016/j.jevs.2026.106064
Borges NF, Barros ML, Silva IS, Barddal JEI, de Freitas AFB, Sumiyoshi MH, Muraro LS, Gomes SSP, de Oliveira ACS, Ribeiro MG, de Aguiar DM.Brucellosis is a chronic infectious disease in equids, characterized by suppurative lesions in joints and bursae, with potential impacts on animal performance and public health. Objective: To investigate the prevalence and spatial distribution of Brucella spp. seropositive equids in the state of Mato Grosso, Brazil. Methods: A total of 3,858 serum samples from 3,230 horses, 602 mules, and 26 donkeys were analyzed using the Buffered Acidified Antigen Test (AAT) and Complement Fixation Test (CFT), following Brazilian Ministry of Agriculture guidelines. Samples were obtained from all municipaliti...
A Breed-Specific SNP Panel With Higher Theoretical Exclusion Power and Practical Equivalence to International Standards for Parentage Testing in Pura Raza Española Horses.
Animal genetics    June 12, 2026   Volume 57, Issue 3 e70147 doi: 10.1002/age.70147
Laseca N, Molina A, Marshall T, González I, Poyato-Bonilla J, Valera M.Accurate parentage assignment is essential for managing livestock breeding and genetic improvement. This study evaluated a breed-specific SNP panel (375 SNPs) for the Pura Raza Española (PRE) horse, comparing its performance against the traditional STR reference panel (18 microsatellite) and the international ISAG SNP core panel (378 SNPs) across population scenarios differing in levels of inbreeding and kinship. The PRE-specific SNPs were selected from the Axiom EQUIGENE array based on high genotyping reliability (call-rate > 99%), Mendelian error minimization, Hardy-Weinberg equilibr...
Current consensus and controversies in equine parasite control.
The Veterinary record    June 12, 2026   Volume 198, Issue 12 e514-e516 doi: 10.1002/vetr.70887
Shrubb J.Julia Shrubb, chair of CANTER - Controlling ANTiparasitic resistance in Equines Responsibly - takes a look at where there is currently agreement on equine parasite control practices and where debate is ongoing.
Prevalence of gastrointestinal parasites in horses in the district of Abidjan, Ivory Coast.
Scientific reports    June 12, 2026   doi: 10.1038/s41598-026-56528-z
Kante N, Kaba L, N'Gotta KI, Camara AOD, Salako AS, Berete K, Tuo K, N'Docho AF, Gondo É, Teohou MMC, N'Gotta AA, Traore AK, Barro N.Gastrointestinal parasitic infections constitute a major constraint to equine health and productivity, particularly in tropical regions where climatic and environmental conditions favor parasite survival and transmission. In Côte d'Ivoire, epidemiological data regarding gastrointestinal parasites in horses remain limited, especially in urban and peri-urban settings such as the district of Abidjan. This study aimed to determine the prevalence, diversity, and associated risk factors of gastrointestinal parasites affecting horses in the study area. A descriptive and analytical cross-sectional st...
Evaluation of a canine Coll2-1 ELISA kit for use in equine serum and its correlation with radiographic joint disease.
Journal of equine veterinary science    June 12, 2026   106063 doi: 10.1016/j.jevs.2026.106063
Ramos S, Lamy E, Nunes T, Rosa T, Bettencourt E, Monteiro S.This study evaluated an enzyme-linked immunosorbent assay (ELISA) for the detection of Coll2-1, a peptide fragment of type II collagen released during cartilage degradation in horses using a kit originally developed for dogs. Objective: The objectives of this study were to: 1) assess the analytical applicability of the canine Coll2-1 ELISA in equine serum and 2) explore the correlations between Coll2-1 concentrations and radiographic scores of osteoarthritis (OA) in horses. Methods: Twelve Lusitano stallions, of similar age and workload but presenting different radiographic scores were include...
An unusual presentation of a sarcoid in Australia.
Australian veterinary journal    June 12, 2026   doi: 10.1111/avj.70104
Swadling VD, Hardcastle MR, Munday JS.Sarcoids are a common skin tumour affecting all equids. To date, only one case of a sarcoid in the oral cavity of the horse has been reported. In the present case, an oral sarcoid was identified using histology and polymerase chain reaction to detect bovine papillomavirus type 2 DNA. The lesion was known to be incompletely surgically excised and did not recur over a 15-month period. This case report describes the first oral sarcoid identified in Australia.
Observations on Anti-Predator Defense Behavior in Feral Horses in Venezuela.
Animals : an open access journal from MDPI    June 12, 2026   Volume 16, Issue 12 1826 doi: 10.3390/ani16121826
Rees L, Kieson E.Apart from vigilance and flight, anti-predator defense behavior in horses has not been well documented despite its importance during natural selection. In this study, observations of a feral herd (around 140) of Venezuelan horses sympatric with puma and jaguar divided such defense into precaution and reaction. Group living and the avoidance of danger areas are precautionary measures enhanced by the stallion's vigilance and his actions to keep small foals with the band. Reactions to perceived threats comprise communication of alarm; bunching, or cohesion, as a primary response; massed flight fo...
Maternal age-related changes in renin-angiotensin-aldosterone system and electrolyte homeostasis during equine pregnancy.
Journal of equine veterinary science    June 12, 2026   Volume 165 106057 doi: 10.1016/j.jevs.2026.106057
Satué K, Fazio E, Domingo R, Velasco-Martínez MG, Cravana C, La Fauci D.The renin-angiotensin-aldosterone system (RAAS) plays a key role in fluid and electrolyte homeostasis during pregnancy. In humans and laboratory animals, maternal age has been shown to modulate RAAS activity, potentially altering the secretion of renin (REN), angiotensin II (ANG-II), and aldosterone (ALD), thereby influencing sodium (Na⁺) and potassium (K⁺) regulation. Objective: To evaluate whether maternal age influences RAAS activity in pregnant mares. Methods: Forty-five Spanish Purebred mares were monitored monthly over 11 months of pregnancy and categorized into three age groups: You...
Skull morphometry and sexual dimorphism in the Kyrgyz horse: A multivariate size-corrected analysis.
Research in veterinary science    June 11, 2026   Volume 210 106288 doi: 10.1016/j.rvsc.2026.106288
Arı HH, Duro S, Alpak H, Tulobaev A, Ceylan N, Ashyk A, Salykov R, Djordjevic M, Gündemir O.This study aimed to evaluate sexual dimorphism in Kyrgyz horse skulls by separating size and shape components using Mosimann log-shape ratios, based on 18 linear cranial measurements recorded from 39 specimens. Raw linear data were analyzed alongside size-corrected Mosimann log-shape ratios to distinguish overall size from size-independent shape. Males showed larger raw means than females for all 18 variables, and 14 measurements remained significant after false-discovery-rate correction. The strongest raw differences involved skull length to nasale, total skull length, nasal length, breadth b...
The “gait keeper” mutation does not influence gait scores in Pêga donkeys: a preliminary result.
Journal of equine veterinary science    June 11, 2026   Volume 165 106056 doi: 10.1016/j.jevs.2026.106056
Santana CJS, Rocha VCP, de Moraes JVS, Senes BB, Maciel SVSA, Oliveira IPP, Costa RB, Alves JS, de Camargo GMF.The DMRT3 "gait keeper" mutation is a major determinant of locomotor patterns in horses, but its role in donkeys remains unclear. This study evaluated the association between this polymorphism and gait performance in Pêga donkeys, a breed selected for gaited locomotion. A total of 260 animals were phenotyped for gait scores and genotyped by PCR-sequencing targeting exon 2 of DMRT3. All individuals were homozygous for the wild-type allele (CC), and the mutant A allele was not detected. The monomorphism precludes any association with gait scores, possibly indicating that this DMRT3 mutation has...
Machine learning-based gait classification and genome-wide association identify a QTL for gait type in Colombian paso horses.
iScience    June 11, 2026   Volume 29, Issue 7 116320 doi: 10.1016/j.isci.2026.116320
Novoa-Bravo M, Meadows JRS, Serra-Bragança F, van de Vall B, Kullander K, Rhodin M, Lindgren G.Coordinated mammalian locomotion relies on spinal circuits where regulates strides and alternative gaits. However, does not explain differences between the Colombian paso horse breed's specialized gaits, Colombian trocha and Colombian trot. We used inertial sensors and machine learning to develop accurate phenotyping ( = 225 horses), before performing genome-wide association analysis ( = 85 horses, 670K array). We identified a 2.43 Mb quantitative trait locus (QTL) on ECA16, where haplotypes featuring lead variants (rs1147402472, = 1.95 × 10; rs1136628503, = 8.52 × 10) explained 48.6% of...
Effect of Antioxidant Supplementation on Systemic Oxidative Status in Breeding Stallions During Intensive Semen Collection.
Animals : an open access journal from MDPI    June 11, 2026   Volume 16, Issue 12 1804 doi: 10.3390/ani16121804
Del Prete C, Ruggiero A, Longobardi C, Cutrignelli MI, Vastolo A.Breeding stallions are subjected to increased metabolic and physiological demands during the reproductive season, which may lead to enhanced production of reactive oxygen species and disruption of redox balance. The present study evaluated the effects of dietary antioxidant supplementation on oxidative status, metabolic profile, and hindgut fermentation in stallions undergoing intensive semen collection activity. Ten adult breeding stallions were assigned either to a control group or to a treatment group receiving a commercial antioxidant supplement (Oxyliver) for 60 days, followed by a 60-day...
Development of an indirect ELISA for the detection of venezuelan equine encephalitis virus specific antibodies in horses.
PloS one    June 10, 2026   Volume 21, Issue 6 e0338819 doi: 10.1371/journal.pone.0338819
Chevrier M, Mathews-Martin L, Mariteragi-Helle T, Dumarest M, Boudjadi Y, Bréard E, Turpaud M, Deshayes T, Vissani MA, Vinueza RL, Coello Peralta R....Venezuelan equine encephalitis virus (VEEV) is a re-emerging zoonotic pathogen to which equines are most susceptible mammals hosts with a high mortality rate raging from19 to 83%. The virus, transmitted between vertebrates and mosquitos, causes unpredictable sporadic epizootic outbreaks in equids and humans across Central and South America. Although VEEV is not currently circulating in Europe the combination of global change, increasing international trade and the presence of potentially competent vectors (mosquitoes of the genera Aedes and Culex), raises a non-negligible risk of its introduct...
A multimodal approach to equine thoracolumbar myofascial pain: integrating clinical examination, AI-based gait analysis, and the ridden horse pain ethogram.
BMC veterinary research    June 9, 2026   doi: 10.1186/s12917-026-05566-w
Resano-Zuazu M, Carmona JU, Argüelles D.Myofascial pain syndrome (MPS) is considered highly prevalent in horses, yet it is frequently underdiagnosed due to limited understanding of its clinical presentation and functional consequences, as well as the inherent challenges associated with its clinical assessment and objective measurement. In equines, the relationship between palpation-based MPS indicators, gait asymmetries, and pain-related behaviours during ridden work remains unclear. The primary aim of this study was to evaluate the association between thoracolumbar palpation pain scores and gait asymmetries measured in-hand and dur...
Cytogenetic, genomic, and reproductive characterization of four pura Raza Española mares carrying a 64, X, del(X),t(X; Y) complex karyotype.
Journal of animal science    June 9, 2026   skag185 doi: 10.1093/jas/skag185
Arjona-Delgado MY, Pirosanto Y, Raudsepp T, Vargas-Pérez MÁ, Valera M, Encina A, Azor P, Bugno-Poniewierska M, Demyda-Peyrás S.Chromosomal abnormalities involving the sex chromosomes constitute critical genetic factors influencing fertility in horses. Nevertheless, a substantial proportion of individuals carrying sex chromosomal abnormalities exhibit normal reproductive phenotypes, which complicates their identification until adulthood. Here, we present four mares carrying a complex deletion/translocation rearrangement with the karyotype formula 64, X, del(X)(p∼25),t(X; Y)(p∼25; q∼15). The cases analyzed belong to three unrelated families of Pura Raza Española (PRE), all presenting the same chromosomal abnormal...
A retrospective study of morbidity and mortality in captive Przewalski’s horses (Equus przewalskii) between 1973 and 2021.
Journal of comparative pathology    June 9, 2026   Volume 228 1-9 doi: 10.1016/j.jcpa.2026.05.004
Quandel CP, Gohl C, Steinmetz HW, Blutke A, Denk D.Przewalski's horses (Equus przewalskii) were declared extinct in the wild in 1996, but reintroduction programmes initiated in 1997 have enabled the re-establishment of free-ranging populations. The maintenance of healthy captive populations remains essential to support these conservation efforts, yet systematic pathological investigations in captive Przewalski's horses are lacking. This study provides the first necropsy-based overview of morbidity and mortality in a captive European population. Pathology reports from 64 horses examined between 1973 and 2021 were reviewed retrospectively. Acros...
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