Analyze Diet

Topic:Sensory Perception

Sensory perception in horses involves the processing and interpretation of sensory stimuli from the environment through various sensory modalities. Horses rely on their senses, including vision, hearing, touch, taste, and smell, to interact with their surroundings and respond to potential threats or opportunities. Vision in horses is adapted for detecting movement and changes in light, with a wide field of view due to the lateral placement of their eyes. Their hearing is sensitive to a range of frequencies, allowing them to detect sounds over long distances. The tactile sense is facilitated by whiskers and sensitive skin, particularly around the muzzle, aiding in exploration and communication. Taste and smell play roles in feeding behavior and social interactions. This page compiles peer-reviewed research studies and scholarly articles that explore the anatomical structures, physiological mechanisms, and behavioral implications of sensory perception in equine species.
Horses (Equus caballus) select the greater of two quantities in small numerical contrasts.
Animal cognition    April 23, 2009   Volume 12, Issue 5 733-738 doi: 10.1007/s10071-009-0225-0
Uller C, Lewis J.The ability to select the greater numerosity over another in small sets seems to stem from the calculation of which set contains more, and has been taken as evidence of a primordial representation at the roots of the primate numerical system. We tested 56 horses (Equus caballus) in a paradigm previously used with human infants and nonhuman primates. Horses saw two quantities paired in contrasts-2 versus 1, 3 versus 2, 6 versus 4 and a control for volume, 2 versus 1 big-and had to make a choice by snout touching the container holding the numerosity selected. The horses spontaneously selected th...
Stimulus discrimination by horses under scotopic conditions.
Behavioural processes    April 21, 2009   Volume 82, Issue 1 45-50 doi: 10.1016/j.beproc.2009.04.009
Hanggi EB, Ingersoll JF.Scotopic vision in horses (Equus caballus) was investigated using behavioral measurements for the first time. Four horses were tested for the ability to make simple visual discriminations of geometric figures (circles and triangles) under various brightness levels within an enclosed building. Measurements of brightness ranging from 10.37 to 24.12 magnitudes per square arcsecond (mag/arcsec(2); in candelas per square meter-7.70 to 2.43E-05cd/m(2)) were taken using a Sky Quality Meter. These values approximated outdoor conditions ranging from twilight in open country to a dark moonless night in ...
How horses (Equus caballus) see the world: humans as significant “objects”.
Animal cognition    April 21, 2009   Volume 12, Issue 4 643-654 doi: 10.1007/s10071-009-0223-2
Fureix C, Jego P, Sankey C, Hausberger M.This study aimed to determine whether horses have a kind of memory of humans (based on previous interactions), leading to a general significance of humans revealed by their reactions to humans in subsequent interactions. Subjects were 59 adult horses used to interact daily with humans. Three types of behavioural tests involving an unknown experimenter evaluated three possibly different memorized types of human-animal interactions (not work-related, using work-related objects, unfamiliar working task). We also performed standardized observations of routine interactions between each horse and it...
Policy Delphi with vignette methodology as a tool to evaluate the perception of equine welfare.
Veterinary journal (London, England : 1997)    April 17, 2009   Volume 181, Issue 1 63-69 doi: 10.1016/j.tvjl.2009.03.012
Collins J, Hanlon A, More SJ, Wall PG, Duggan V.A three-round Policy Delphi using vignette methodology was employed as a new approach to study stakeholder perceptions and experiences of equine welfare. Forty-four representatives from stakeholder groups in the Irish equine industry participated. In Round 1, vignettes (narratives illustrating potential infringements of equine welfare) were presented to assess perceptions of 'Acceptability' and experiences of 'Frequency of occurrence'. In Round 2, lists of situations where equine welfare might be compromised, possible drivers of behaviour and potential solutions were presented for grading. In ...
The relationship between visual memory and rider expertise in a show-jumping context.
Veterinary journal (London, England : 1997)    April 16, 2009   Volume 181, Issue 1 29-33 doi: 10.1016/j.tvjl.2009.03.007
Hall C, Liley C, Murphy J, Crundall D.Individuals develop visual skills whilst participating in sport. The aim of this study was to assess the effect of reported riding expertise on the recall of visual information relating to show-jumps. Relevant (F1) and irrelevant (F2) points of focus were identified in 22 photographs of show-jumps. Participants were students (n=40) with varying levels of horse-riding ability. After viewing each photograph for 4s, the task was to identify F1s or F2s from four alternatives viewed for 10s. F1s were recalled significantly more than F2s (P<0.001). Riding expertise did not affect overall recall but ...
Long-term memory for categories and concepts in horses (Equus caballus).
Animal cognition    January 16, 2009   Volume 12, Issue 3 451-462 doi: 10.1007/s10071-008-0205-9
Hanggi EB, Ingersoll JF.Three horses (Equus caballus) with a history of performing cognitive tasks including discrimination learning, categorization, and concept use were tested to evaluate their long-term memory (LTM) in three experiments. In addition, use of LCD multi-displays for stimulus presentation was incorporated into cognition testing protocol for the first time with horses. Experiment 1 tested LTM for discrimination learning that originally occurred 6 years earlier. Five sets of stimuli were used and the two horses tested showed no decrement in performance on four of the sets; however, both horses did score...
Cross-modal individual recognition in domestic horses (Equus caballus).
Proceedings of the National Academy of Sciences of the United States of America    December 15, 2008   Volume 106, Issue 3 947-951 doi: 10.1073/pnas.0809127105
Proops L, McComb K, Reby D.Individual recognition is considered a complex process and, although it is believed to be widespread across animal taxa, the cognitive mechanisms underlying this ability are poorly understood. An essential feature of individual recognition in humans is that it is cross-modal, allowing the matching of current sensory cues to identity with stored information about that specific individual from other modalities. Here, we use a cross-modal expectancy violation paradigm to provide a clear and systematic demonstration of cross-modal individual recognition in a nonhuman animal: the domestic horse. Su...
Position of the head is not associated with changes in horse vision.
Equine veterinary journal    November 26, 2008   Volume 40, Issue 6 599-601 doi: 10.2746/042516408x332417
Bartos L, Bartosová J, Starostová L.It has become accepted that the horse cannot see directly in front when the nose is lowered and must therefore rely on the rider. We tested the hypothesis that this conclusion would be correct only if the horse did not adjust the eyeball horizontal axis to changes of the head position. The results of the present study suggest that it is unlikely that horses have limited vision in relation to their head position when driven by the rider, and that the horse maintains the optimal horizontal eyeball position regardless of head position relative to the ground.
Hind limb stepping over obstacles in the horse guided by place-object memory.
Behavioural brain research    November 24, 2008   Volume 198, Issue 2 372-379 doi: 10.1016/j.bbr.2008.11.023
Whishaw IQ, Sacrey LA, Gorny B.An animal that has stepped over an obstacle with its forelimbs uses a memory of the obstacle to guide the hind limbs so that they also clear the obstacle, even in situations in which long pauses are introduced between forelimb and hind limb stepping. To further clarify the features of hind limb obstacle clearance memory, the present study examined hind limb obstacle clearance in the horse. A rider guided horses over obstacles and paused the horse over obstacles in tests that examined the relationship between forelimb and hind limb stepping, with the following results. First, the horses display...
The absolute threshold of colour vision in the horse.
PloS one    November 12, 2008   Volume 3, Issue 11 e3711 doi: 10.1371/journal.pone.0003711
Roth LS, Balkenius A, Kelber A.Arrhythmic mammals are active both during day and night if they are allowed. The arrhythmic horses are in possession of one of the largest terrestrial animal eyes and the purpose of this study is to reveal whether their eye is sensitive enough to see colours at night. During the day horses are known to have dichromatic colour vision. To disclose whether they can discriminate colours in dim light a behavioural dual choice experiment was performed. We started the training and testing at daylight intensities and the horses continued to choose correctly at a high frequency down to light intensitie...
Laterality and emotions: visual laterality in the domestic horse (Equus caballus) differs with objects’ emotional value.
Physiology & behavior    March 12, 2008   Volume 94, Issue 3 487-490 doi: 10.1016/j.physbeh.2008.03.002
De Boyer Des Roches A, Richard-Yris MA, Henry S, Ezzaouïa M, Hausberger M.Lateralization of emotions has received great attention in the last decades, both in humans and animals, but little interest has been given to side bias in perceptual processing. Here, we investigated the influence of the emotional valence of stimuli on visual and olfactory explorations by horses, a large mammalian species with two large monocular visual fields and almost complete decussation of optic fibres. We confronted 38 Arab mares to three objects with either a positive, negative or neutral emotional valence (novel object). The results revealed a gradient of exploration of the 3 objects ...
An investigation of colour discrimination with horses (Equus caballus).
Behavioural processes    February 13, 2008   Volume 78, Issue 3 387-396 doi: 10.1016/j.beproc.2008.02.003
Blackmore TL, Foster TM, Sumpter CE, Temple W.The ability of four horses (Equus caballus) to discriminate coloured (three shades of blue, green, red, and yellow) from grey (neutral density) stimuli, produced by back projected lighting filters, was investigated in a two response forced-choice procedure. Pushes of the lever in front of a coloured screen were occasionally reinforced, pushes of the lever in front of a grey screen were never reinforced. Each colour shade was randomly paired with a grey that was brighter, one that was dimmer, and one that approximately matched the colour in terms of brightness. Each horse experienced the colour...
Comprehension of human pointing gestures in horses (Equus caballus).
Animal cognition    February 5, 2008   Volume 11, Issue 3 457-466 doi: 10.1007/s10071-008-0136-5
Maros K, Gácsi M, Miklósi A.Twenty domestic horses (Equus caballus) were tested for their ability to rely on different human gesticular cues in a two-way object choice task. An experimenter hid food under one of two bowls and after baiting, indicated the location of the food to the subjects by using one of four different cues. Horses could locate the hidden reward on the basis of the distal dynamic-sustained, proximal momentary and proximal dynamic-sustained pointing gestures but failed to perform above chance level when the experimenter performed a distal momentary pointing gesture. The results revealed that horses coul...
Asymmetry of flight and escape turning responses in horses.
Laterality    August 23, 2007   Volume 12, Issue 5 464-474 doi: 10.1080/13576500701495307
Austin NP, Rogers LJ.We investigated whether horses display greater reactivity to a novel stimulus presented in the left compared to the right monocular visual field, and whether a population bias exists for escape turning when the same stimulus was presented binocularly. Domestic horses (N=30) were tested on three occasions by a person opening an umbrella five metres away and then approaching. The distance each horse moved away before stopping was measured. Distance was greatest for approach on the left side, indicating right hemisphere control of flight behaviour, and thus followed the same pattern found previou...
Colour perception in a dichromat.
The Journal of experimental biology    August 11, 2007   Volume 210, Issue Pt 16 2795-2800 doi: 10.1242/jeb.007377
Roth LS, Balkenius A, Kelber A.Most mammals have dichromatic colour vision based on two different types of cones: a short-wavelength-sensitive cone and a long-wavelength-sensitive cone. Comparing the signal from two cone types gives rise to a one-dimensional chromatic space when brightness is excluded. The so-called ;neutral point' refers to the wavelength that the animal cannot distinguish from achromatic light such as white or grey because it stimulates both cone types equally. The question is: how do dichromats perceive their chromatic space? Do they experience a continuous scale of colours or does the neutral point divi...
[Measurement of the attention time in the horse].
Schweizer Archiv fur Tierheilkunde    March 9, 2007   Volume 149, Issue 2 77-83 doi: 10.1024/0036-7281.149.2.77
Rapin V, Poncet PA, Burger D, Mermod C, Richard MA.A study carried out on 49 horses showed that it is possible to measure the attention time by operant conditioning. After teaching horses an instrumental task using a signal, we were then able to test their attention time by asking them to prolong it increasingly while setting success and failure criteria. Two tests were performed 3 weeks apart. The 2nd test was feasible without relearning, a proof of memory, and was repeatable, a proof of consistency in the attention time. A significant difference was observed between the 3 age groups. Young horses often performed very well during the 1st test...
Social learning in horses from a novel perspective.
Behavioural processes    March 6, 2007   Volume 76, Issue 1 37-60 doi: 10.1016/j.beproc.2006.08.010
Krueger K, Flauger B.No abstract available
The impact of visual perception on equine learning.
Behavioural processes    March 4, 2007   Volume 76, Issue 1 29-60 doi: 10.1016/j.beproc.2006.09.017
Hall C.No abstract available
Imprinting training and conditioned taste aversion.
Behavioural processes    March 4, 2007   Volume 76, Issue 1 14-60 doi: 10.1016/j.beproc.2006.09.016
Houpt KA.No abstract available
Equine learning behaviour.
Behavioural processes    March 4, 2007   Volume 76, Issue 1 1-13 doi: 10.1016/j.beproc.2006.06.009
Murphy J, Arkins S.Scientists and equestrians continually seek to achieve a clearer understanding of equine learning behaviour and its implications for training. Behavioural and learning processes in the horse are likely to influence not only equine athletic success but also the usefulness of the horse as a domesticated species. However given the status and commercial importance of the animal, equine learning behaviour has received only limited investigation. Indeed most experimental studies on equine cognitive function to date have addressed behaviour, learning and conceptualization processes at a moderately ba...
Color vision in horses (Equus caballus): deficiencies identified using a pseudoisochromatic plate test.
Journal of comparative psychology (Washington, D.C. : 1983)    February 28, 2007   Volume 121, Issue 1 65-72 doi: 10.1037/0735-7036.121.1.65
Hanggi EB, Ingersoll JF, Waggoner TL.In the past, equine color vision was tested with stimuli composed either of painted cards or photographic slides or through physiological testing using electroretinogram flicker photometry. Some studies produced similar results, but others did not, demonstrating that there was not yet a definitive answer regarding color vision in horses (Equus caballus). In this study, a pseudoisochromatic plate test--which is highly effective in testing color vision both in small children and in adult humans--was used for the first time on a nonhuman animal. Stimuli consisted of different colored dotted circl...
Cone excitation ratios correlate with color discrimination performance in the horse (Equus caballus).
Journal of comparative psychology (Washington, D.C. : 1983)    November 23, 2006   Volume 120, Issue 4 438-448 doi: 10.1037/0735-7036.120.4.438
Hall CA, Cassaday HJ, Vincent CJ, Derrington AM.Six horses (Equus caballus) were trained to discriminate color from grays in a counterbalanced sequence in which lightness cues were irrelevant. Subsequently, the pretrained colors were presented in a different sequence. Two sets of novel colors paired with novel grays were also tested. Performance was just as good in these transfer tests. Once the horse had learned to select the chromatic from the achromatic stimulus, regardless of the specific color, they were immediately able to apply this rule to novel stimuli. In terms of the underlying visual mechanisms, the present study showed for the ...
Influence of various early human-foal interferences on subsequent human-foal relationship.
Developmental psychobiology    November 18, 2006   Volume 48, Issue 8 712-718 doi: 10.1002/dev.20189
Henry S, Richard-Yris MA, Hausberger M.Whereas the way animals perceive human contact has been particularly examined in pet animals, a small amount of investigations has been done in domestic ungulates. It was nevertheless assumed that, as pet animals, non-aggressive forms of tactile contact were as well rewarding or positive for these species, even though the features of intraspecific relationships in pet animals and domestic ungulates may be to some extent different. We test here the hypothesis that horses may not consider physical handling by humans as a positive event. When comparing different early human-foal interactions, we ...
Evidence of bias affecting the interpretation of the results of local anaesthetic nerve blocks when assessing lameness in horses.
The Veterinary record    September 12, 2006   Volume 159, Issue 11 346-349 doi: 10.1136/vr.159.11.346
Arkell M, Archer RM, Guitian FJ, May SA.Eighteen observers were influenced to different extents in the grades of lameness they allocated to eight horses by whether they knew that a nerve block had been administered; on a scale from 0 to 10 the mean difference in grade allocated once the observer knew a horse had been nerve-blocked was increased by 0.4. The consistency of the assessments made by the individual observers was good, with a an average of 0.6 of a grade difference when grading the same horse on two occasions. The agreement between the assessments of four orthopaedic experts was reasonable (+/-1 grade), but significantly p...
Laterality of horses associated with emotionality in novel situations.
Laterality    June 7, 2006   Volume 11, Issue 4 355-367 doi: 10.1080/13576500600624221
Larose C, Richard-Yris MA, Hausberger M, Rogers LJ.We have established that lateral biases are characteristic of visual behaviour in 65 horses. Two breeds, Trotters and French Saddlebreds aged 2 to 3, were tested on a novel object test. The main finding was a significant correlation between emotionality index and the eye preferred to view the novel stimulus: the higher the emotionality, the more likely that the horse looked with its left eye. The less emotive French Saddlebreds, however, tended to glance at the object using the right eye, a tendency that was not found in the Trotters, although the emotive index was the same for both breeds. Th...
Perceptions of animal physiotherapy amongst Irish veterinary surgeons.
Irish veterinary journal    February 1, 2006   Volume 59, Issue 2 85-89 doi: 10.1186/2046-0481-59-2-85
Doyle A, Horgan NF.The aim of this study was to investigate veterinary surgeons' perceptions, knowledge and use of animal physiotherapy in the Republic of Ireland. A questionnaire was developed and sent to 200 veterinary surgeons, of which 97 were returned. Results indicated that 77 (79%) of respondents were aware of animal physiotherapists. Common sources of information included veterinary colleagues, owners and professional journals, with physiotherapists themselves and undergraduate training being less commonly cited. Awareness of animal physiotherapy was greatest amongst those working in equine practice (χ2...
Effects of a therapeutic riding program on at-risk and special education children.
Journal of the American Veterinary Medical Association    January 24, 2006   Volume 228, Issue 1 46-52 doi: 10.2460/javma.228.1.46
Kaiser L, Smith KA, Heleski CR, Spence LJ.To determine the effects of a therapeutic riding program on psychosocial measurements among children considered at risk for poor performance or failure in school or life and among children in special education programs. Methods: Observational study. Methods: 17 at-risk children (6 boys and 11 girls) and 14 special education children (7 boys and 7 girls). Methods: For the at-risk children, anger, anxiety, perceived self-competence, and physical coordination were assessed. For the special education children, anger and cheerfulness were measured, and the children's and their mothers' perceptions ...
Double blind, you are the weakest link–goodbye!
Equine veterinary journal    November 22, 2005   Volume 37, Issue 6 557-558 doi: 10.2746/042516405775314916
Devereaux PJ, Bhandari M, Montori VM, Manns BJ, Ghali WA, Guyatt GH.No abstract available
Frontier medicine: the future and integrative medicine.
The Veterinary clinics of North America. Equine practice    January 22, 2005   Volume 17, Issue 2 351-377 doi: 10.1016/s0749-0739(17)30067-6
Ross C, Haussler KK, Kenney JD, Marks D, Bertone JJ, Henneman K, May KJ.Vigorous and prolonged effort is required to gain true mastery of the healing arts. Conventional and complementary medicine have complementary strengths and weaknesses. Like the yin and yang of traditional Chinese medicine, they naturally flow into one another by a process of induction, creating balance. Integrative medicine is the frontier; it is the future. If we are to progress beyond our current understanding and ability to heal, we must work with theoretic models that allow us and our perception to operate "outside the box." For some, this understanding is intuitive. It is through coopera...
Hippotherapy.
Physical medicine and rehabilitation clinics of North America    October 2, 2004   Volume 15, Issue 4 843-vii doi: 10.1016/j.pmr.2004.02.002
Meregillano G.Hippotherapy refers to the use of the movement of the horse asa treatment tool by physical therapists, occupational therapists,and speech-language therapists to address impairments, functional limitations, and disabilities in clients with neuromusculoskeletal dysfunction, such as cerebral palsy. Hippotherapy is used as part of an integrated treatment program to achieve functional outcomes. Hippotherapy engages the client in activities on the horse that are enjoyable and challenging. In the controlled hippotherapyenvironment, the therapist modifies the horse's movement and carefully grades sens...